HRP20120294T1 - Suncokretove biljke otporne na herbicide, za polinukleotide koji kodiraju velike pojedinačne proteine otporne na herbicide sintezom acetohidroksilne kiseline, te postupci uporabe - Google Patents
Suncokretove biljke otporne na herbicide, za polinukleotide koji kodiraju velike pojedinačne proteine otporne na herbicide sintezom acetohidroksilne kiseline, te postupci uporabe Download PDFInfo
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- HRP20120294T1 HRP20120294T1 HRP20120294TT HRP20120294T HRP20120294T1 HR P20120294 T1 HRP20120294 T1 HR P20120294T1 HR P20120294T T HRP20120294T T HR P20120294TT HR P20120294 T HRP20120294 T HR P20120294T HR P20120294 T1 HRP20120294 T1 HR P20120294T1
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- methyl
- herbicide
- imidazolinone
- herbicides
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- 239000004009 herbicide Substances 0.000 title claims abstract 27
- 241000208818 Helianthus Species 0.000 title claims abstract 17
- 230000002363 herbicidal effect Effects 0.000 title claims abstract 16
- 239000002157 polynucleotide Substances 0.000 title claims abstract 7
- 102000040430 polynucleotide Human genes 0.000 title claims abstract 7
- 108091033319 polynucleotide Proteins 0.000 title claims abstract 7
- 238000000034 method Methods 0.000 title claims 6
- 101710103719 Acetolactate synthase large subunit Proteins 0.000 title 1
- 101710182467 Acetolactate synthase large subunit IlvB1 Proteins 0.000 title 1
- 101710171176 Acetolactate synthase large subunit IlvG Proteins 0.000 title 1
- 101710196435 Probable acetolactate synthase large subunit Proteins 0.000 title 1
- CAAMSDWKXXPUJR-UHFFFAOYSA-N 3,5-dihydro-4H-imidazol-4-one Chemical compound O=C1CNC=N1 CAAMSDWKXXPUJR-UHFFFAOYSA-N 0.000 claims abstract 13
- 241000196324 Embryophyta Species 0.000 claims abstract 10
- 108090000623 proteins and genes Proteins 0.000 claims abstract 5
- 102000004169 proteins and genes Human genes 0.000 claims abstract 3
- 239000002253 acid Substances 0.000 claims abstract 2
- 230000015572 biosynthetic process Effects 0.000 claims abstract 2
- 238000003786 synthesis reaction Methods 0.000 claims abstract 2
- 239000000203 mixture Substances 0.000 claims 6
- 229940100389 Sulfonylurea Drugs 0.000 claims 5
- 150000001413 amino acids Chemical group 0.000 claims 5
- YROXIXLRRCOBKF-UHFFFAOYSA-N sulfonylurea Chemical class OC(=N)N=S(=O)=O YROXIXLRRCOBKF-UHFFFAOYSA-N 0.000 claims 5
- YWBFPKPWMSWWEA-UHFFFAOYSA-O triazolopyrimidine Chemical compound BrC1=CC=CC(C=2N=C3N=CN[N+]3=C(NCC=3C=CN=CC=3)C=2)=C1 YWBFPKPWMSWWEA-UHFFFAOYSA-O 0.000 claims 4
- 235000001014 amino acid Nutrition 0.000 claims 2
- 230000002401 inhibitory effect Effects 0.000 claims 2
- 235000018102 proteins Nutrition 0.000 claims 2
- NUPJIGQFXCQJBK-UHFFFAOYSA-N 2-(4-isopropyl-4-methyl-5-oxo-4,5-dihydro-1H-imidazol-2-yl)-5-(methoxymethyl)nicotinic acid Chemical compound OC(=O)C1=CC(COC)=CN=C1C1=NC(C)(C(C)C)C(=O)N1 NUPJIGQFXCQJBK-UHFFFAOYSA-N 0.000 claims 1
- CLQMBPJKHLGMQK-UHFFFAOYSA-N 2-(4-isopropyl-4-methyl-5-oxo-4,5-dihydro-1H-imidazol-2-yl)nicotinic acid Chemical compound N1C(=O)C(C(C)C)(C)N=C1C1=NC=CC=C1C(O)=O CLQMBPJKHLGMQK-UHFFFAOYSA-N 0.000 claims 1
- CABMTIJINOIHOD-UHFFFAOYSA-N 2-[4-methyl-5-oxo-4-(propan-2-yl)-4,5-dihydro-1H-imidazol-2-yl]quinoline-3-carboxylic acid Chemical compound N1C(=O)C(C(C)C)(C)N=C1C1=NC2=CC=CC=C2C=C1C(O)=O CABMTIJINOIHOD-UHFFFAOYSA-N 0.000 claims 1
- PVSGXWMWNRGTKE-UHFFFAOYSA-N 5-methyl-2-[4-methyl-5-oxo-4-(propan-2-yl)-4,5-dihydro-1H-imidazol-2-yl]pyridine-3-carboxylic acid Chemical compound N1C(=O)C(C(C)C)(C)N=C1C1=NC=C(C)C=C1C(O)=O PVSGXWMWNRGTKE-UHFFFAOYSA-N 0.000 claims 1
- 239000003666 Amidosulfuron Substances 0.000 claims 1
- CTTHWASMBLQOFR-UHFFFAOYSA-N Amidosulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)N(C)S(C)(=O)=O)=N1 CTTHWASMBLQOFR-UHFFFAOYSA-N 0.000 claims 1
- DCXYFEDJOCDNAF-UHFFFAOYSA-N Asparagine Natural products OC(=O)C(N)CC(N)=O DCXYFEDJOCDNAF-UHFFFAOYSA-N 0.000 claims 1
- 239000005472 Bensulfuron methyl Substances 0.000 claims 1
- 239000005514 Flazasulfuron Substances 0.000 claims 1
- HWATZEJQIXKWQS-UHFFFAOYSA-N Flazasulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)C=2C(=CC=CN=2)C(F)(F)F)=N1 HWATZEJQIXKWQS-UHFFFAOYSA-N 0.000 claims 1
- LXKOADMMGWXPJQ-UHFFFAOYSA-N Halosulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)C=2N(N=C(Cl)C=2C(O)=O)C)=N1 LXKOADMMGWXPJQ-UHFFFAOYSA-N 0.000 claims 1
- 239000005567 Imazosulfuron Substances 0.000 claims 1
- NAGRVUXEKKZNHT-UHFFFAOYSA-N Imazosulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)C=2N3C=CC=CC3=NC=2Cl)=N1 NAGRVUXEKKZNHT-UHFFFAOYSA-N 0.000 claims 1
- DCXYFEDJOCDNAF-REOHCLBHSA-N L-asparagine Chemical compound OC(=O)[C@@H](N)CC(N)=O DCXYFEDJOCDNAF-REOHCLBHSA-N 0.000 claims 1
- ROHFNLRQFUQHCH-YFKPBYRVSA-N L-leucine Chemical compound CC(C)C[C@H](N)C(O)=O ROHFNLRQFUQHCH-YFKPBYRVSA-N 0.000 claims 1
- COLNVLDHVKWLRT-QMMMGPOBSA-N L-phenylalanine Chemical compound OC(=O)[C@@H](N)CC1=CC=CC=C1 COLNVLDHVKWLRT-QMMMGPOBSA-N 0.000 claims 1
- AYFVYJQAPQTCCC-GBXIJSLDSA-N L-threonine Chemical compound C[C@@H](O)[C@H](N)C(O)=O AYFVYJQAPQTCCC-GBXIJSLDSA-N 0.000 claims 1
- KZSNJWFQEVHDMF-BYPYZUCNSA-N L-valine Chemical compound CC(C)[C@H](N)C(O)=O KZSNJWFQEVHDMF-BYPYZUCNSA-N 0.000 claims 1
- ROHFNLRQFUQHCH-UHFFFAOYSA-N Leucine Natural products CC(C)CC(N)C(O)=O ROHFNLRQFUQHCH-UHFFFAOYSA-N 0.000 claims 1
- 239000005584 Metsulfuron-methyl Substances 0.000 claims 1
- 239000005586 Nicosulfuron Substances 0.000 claims 1
- BGNQYGRXEXDAIQ-UHFFFAOYSA-N Pyrazosulfuron-ethyl Chemical group C1=NN(C)C(S(=O)(=O)NC(=O)NC=2N=C(OC)C=C(OC)N=2)=C1C(=O)OCC BGNQYGRXEXDAIQ-UHFFFAOYSA-N 0.000 claims 1
- 239000005616 Rimsulfuron Substances 0.000 claims 1
- 239000005623 Thifensulfuron-methyl Substances 0.000 claims 1
- AYFVYJQAPQTCCC-UHFFFAOYSA-N Threonine Natural products CC(O)C(N)C(O)=O AYFVYJQAPQTCCC-UHFFFAOYSA-N 0.000 claims 1
- 239000004473 Threonine Substances 0.000 claims 1
- KZSNJWFQEVHDMF-UHFFFAOYSA-N Valine Natural products CC(C)C(N)C(O)=O KZSNJWFQEVHDMF-UHFFFAOYSA-N 0.000 claims 1
- 229960001230 asparagine Drugs 0.000 claims 1
- 235000009582 asparagine Nutrition 0.000 claims 1
- XMQFTWRPUQYINF-UHFFFAOYSA-N bensulfuron-methyl Chemical group COC(=O)C1=CC=CC=C1CS(=O)(=O)NC(=O)NC1=NC(OC)=CC(OC)=N1 XMQFTWRPUQYINF-UHFFFAOYSA-N 0.000 claims 1
- NSWAMPCUPHPTTC-UHFFFAOYSA-N chlorimuron-ethyl Chemical group CCOC(=O)C1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(Cl)=CC(OC)=N1 NSWAMPCUPHPTTC-UHFFFAOYSA-N 0.000 claims 1
- VJYIFXVZLXQVHO-UHFFFAOYSA-N chlorsulfuron Chemical compound COC1=NC(C)=NC(NC(=O)NS(=O)(=O)C=2C(=CC=CC=2)Cl)=N1 VJYIFXVZLXQVHO-UHFFFAOYSA-N 0.000 claims 1
- 238000010353 genetic engineering Methods 0.000 claims 1
- 230000035784 germination Effects 0.000 claims 1
- FFCCBBNQPIMUJI-UHFFFAOYSA-N methyl 2-(4-isopropyl-4-methyl-5-oxo-2-imidazolin-2-yl)-para-toluate Chemical compound COC(=O)C1=CC=C(C)C=C1C1=NC(C)(C(C)C)C(=O)N1 FFCCBBNQPIMUJI-UHFFFAOYSA-N 0.000 claims 1
- TYIHVCIQMMTVHE-UHFFFAOYSA-N methyl 2-(4-isopropyl-4-methyl-5-oxo-4,5-dihydro-1H-imidazol-2-yl)-5-methylbenzoate Chemical compound COC(=O)C1=CC(C)=CC=C1C1=NC(C)(C(C)C)C(=O)N1 TYIHVCIQMMTVHE-UHFFFAOYSA-N 0.000 claims 1
- ZTYVMAQSHCZXLF-UHFFFAOYSA-N methyl 2-[[4,6-bis(difluoromethoxy)pyrimidin-2-yl]carbamoylsulfamoyl]benzoate Chemical group COC(=O)C1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(OC(F)F)=CC(OC(F)F)=N1 ZTYVMAQSHCZXLF-UHFFFAOYSA-N 0.000 claims 1
- 125000002496 methyl group Chemical group [H]C([H])([H])* 0.000 claims 1
- RSMUVYRMZCOLBH-UHFFFAOYSA-N metsulfuron methyl Chemical group COC(=O)C1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(C)=NC(OC)=N1 RSMUVYRMZCOLBH-UHFFFAOYSA-N 0.000 claims 1
- RTCOGUMHFFWOJV-UHFFFAOYSA-N nicosulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)C=2C(=CC=CN=2)C(=O)N(C)C)=N1 RTCOGUMHFFWOJV-UHFFFAOYSA-N 0.000 claims 1
- 229960003512 nicotinic acid Drugs 0.000 claims 1
- 239000011664 nicotinic acid Substances 0.000 claims 1
- 239000002773 nucleotide Substances 0.000 claims 1
- 125000003729 nucleotide group Chemical group 0.000 claims 1
- COLNVLDHVKWLRT-UHFFFAOYSA-N phenylalanine Natural products OC(=O)C(N)CC1=CC=CC=C1 COLNVLDHVKWLRT-UHFFFAOYSA-N 0.000 claims 1
- MEFOUWRMVYJCQC-UHFFFAOYSA-N rimsulfuron Chemical compound CCS(=O)(=O)C1=CC=CN=C1S(=O)(=O)NC(=O)NC1=NC(OC)=CC(OC)=N1 MEFOUWRMVYJCQC-UHFFFAOYSA-N 0.000 claims 1
- 238000009331 sowing Methods 0.000 claims 1
- ZDXMLEQEMNLCQG-UHFFFAOYSA-N sulfometuron methyl Chemical group COC(=O)C1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(C)=CC(C)=N1 ZDXMLEQEMNLCQG-UHFFFAOYSA-N 0.000 claims 1
- AHTPATJNIAFOLR-UHFFFAOYSA-N thifensulfuron-methyl Chemical group S1C=CC(S(=O)(=O)NC(=O)NC=2N=C(OC)N=C(C)N=2)=C1C(=O)OC AHTPATJNIAFOLR-UHFFFAOYSA-N 0.000 claims 1
- XOPFESVZMSQIKC-UHFFFAOYSA-N triasulfuron Chemical compound COC1=NC(C)=NC(NC(=O)NS(=O)(=O)C=2C(=CC=CC=2)OCCCl)=N1 XOPFESVZMSQIKC-UHFFFAOYSA-N 0.000 claims 1
- YMXOXAPKZDWXLY-QWRGUYRKSA-N tribenuron methyl Chemical group COC(=O)[C@H]1CCCC[C@@H]1S(=O)(=O)NC(=O)N(C)C1=NC(C)=NC(OC)=N1 YMXOXAPKZDWXLY-QWRGUYRKSA-N 0.000 claims 1
- IMEVJVISCHQJRM-UHFFFAOYSA-N triflusulfuron-methyl Chemical group COC(=O)C1=CC=CC(C)=C1S(=O)(=O)NC(=O)NC1=NC(OCC(F)(F)F)=NC(N(C)C)=N1 IMEVJVISCHQJRM-UHFFFAOYSA-N 0.000 claims 1
- 239000004474 valine Substances 0.000 claims 1
- FWMNVWWHGCHHJJ-SKKKGAJSSA-N 4-amino-1-[(2r)-6-amino-2-[[(2r)-2-[[(2r)-2-[[(2r)-2-amino-3-phenylpropanoyl]amino]-3-phenylpropanoyl]amino]-4-methylpentanoyl]amino]hexanoyl]piperidine-4-carboxylic acid Chemical compound C([C@H](C(=O)N[C@H](CC(C)C)C(=O)N[C@H](CCCCN)C(=O)N1CCC(N)(CC1)C(O)=O)NC(=O)[C@H](N)CC=1C=CC=CC=1)C1=CC=CC=C1 FWMNVWWHGCHHJJ-SKKKGAJSSA-N 0.000 abstract 1
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Abstract
Biljka suncokreta, naznačena time, da u svojem genomu sadrži barem jednu kopiju od barem jednog endogenog gena koji sadrži veliku podjedinicu polinukleotida sinteze acetohidroksilne kiseline (AHASL), gdje spomenuti AHASL polinukleotid kodira imidazolinon AHASL1 protein otporan na herbicide, koji sadrži sekvencu amino kiseline iznesenu u SEQ ID NO:2; ili njezina biljka potomak, pri čemu spomenuta biljka suncokreta ili njezin potomak, ima povećanu otpornost na barem jedan imidazolinonski herbicid u usporedbi s divljim tipom biljke suncokreta. Patent sadrži još 11 patentnih zahtjeva.
Claims (12)
1. Biljka suncokreta, naznačena time, da u svojem genomu sadrži barem jednu kopiju od barem jednog endogenog gena koji sadrži veliku podjedinicu polinukleotida sinteze acetohidroksilne kiseline (AHASL), gdje spomenuti AHASL polinukleotid kodira imidazolinon AHASL1 protein otporan na herbicide, koji sadrži sekvencu amino kiseline iznesenu u SEQ ID NO:2; ili njezina biljka potomak, pri čemu spomenuta biljka suncokreta ili njezin potomak, ima povećanu otpornost na barem jedan imidazolinonski herbicid u usporedbi s divljim tipom biljke suncokreta.
2. Biljka suncokreta prema zahtjevu 1, naznačena time, da spomenuta biljka ima pojačanu otpornost na barem jedan herbicid odabran iz skupine koju čine herbicidi sulfoniluree i triazolopirimidinski herbicidi.
3. Biljka suncokreta prema zahtjevu 1 ili 2, naznačena time, da spomenuti AHASL polinukleotid otporan na herbicide sadrži sekvencu nukleotida iznesenu u SEQ ID NO:1.
4. Biljka suncokreta prema bilo kojem od zahtjeva 1 do 3, naznačena time, da spomenuti AHASL protein otporan na herbicide nadalje sadrži barem jedan član odabran iz skupine koju čine:
a) leukin na poziciji amino kiseline 559 ili na ekvivalentnoj poziciji u odnosu na sekvencu amino kiseline iznesenu u SEQ ID NO:12; i
b) bilo koji od asparagina, treonina, fenilalanina ili valina na poziciji amino kiseline 638 ili na ekvivalentnoj poziciji u odnosu na sekvencu amino kiseline iznesenu u SEQ ID NO:12.
5. Sjeme biljke suncokreta prema bilo kojem od zahtjeva 1 do 4, naznačeno time, da spomenuto sjeme sadrži u svojem genomu barem jednu kopiju spomenutog AHASL polinukleotida.
6. Biljka suncokreta prema bilo kojem od zahtjeva 1 do 5, naznačena time, da obuhvaća svojstva imidazolinona otpornog na herbicide linije GM40 ili GM1606, uzorak sjemena linije koja je bila deponirana pri ATCC Patent Deposit Number PTA-6716 odnosno pri ATCC Patent Deposit Number PTA-7606, pri čemu je biljka:
a) potomak linije GM40 ili GM1606;
b) mutant, rekombinant ili derivat genetičkog inženjeringa linije GM40 ili GM1606; ili
c) biljka koja je potomak od barem jedne biljke iz (a) ili (b).
7. Sjeme biljke suncokreta prema zahtjevu 6, naznačeno time, da spomenuto sjeme sadrži svojstva imidazolinona otpornog na herbicide iz linije GM40 ili GM1606.
8. Postupak suzbijanja korova u blizini biljke suncokreta, naznačen time, da spomenuti postupak obuhvaća primjenu učinkovite količine imidazolinon-herbicida, ili mješavine imidazolinon-herbicida, sulfonilurea-herbicida i/ili triazolopirimidin-herbicida, na korov i na biljku suncokreta, pri čemu je spomenuta biljka suncokreta ona biljka suncokreta prema bilo kojem od zahtjeva 1 do 4 ili 6.
9. Postupak prema zahtjevu 8, naznačen time, da je spomenuti imidazolinon-herbicid odabran iz skupine koju čine: 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-nikotinska kiselina, 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-3-kinolinkarboksilna kiselina, 5-etil-2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-nikotinska kiselina, 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-5-(metoksimetil)-nikotinska kiselina, 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-5-metilnikotinska kiselina i mješavina metil-6-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-m-toluata i metil-2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-p-toluata, te njihove mješavine.
10. Postupak prema zahtjevu 8, naznačen time, da je spomenuti sulfonilurea-herbicid odabran iz skupine koju čine: klorosulfuron, metsulfuron metil, sulfometuron metil, klorimuron etil, tifensulfuron metil, tribenuron metil, bensulfuron metil, nikosulfuron, etametsulfuron metil, rimsulfuron, triflusulfuron metil, triasulfuron, primisulfuron metil, kinosulfuron, amidosulfuron, flazasulfuron, imazosulfuron, pirazosulfuron etil, halosulfuron, te njihove mješavine.
11. Sjeme biljke prema bilo kojem od zahtjeva 1 do 4 ili 6, naznačeno time, da se spomenuto sjeme tretira sa AHAS-inhibirajućim imidazolinon-herbicidom, ili mješavinom imidazolinon-herbicida, sulfonilurea-herbicida i/ili triazolopirimidin-herbicida.
12. Postupak za suzbijanje neželjene vegetacije, naznačen time, da obuhvaća stavljanje u doticaj sjemena biljke prema bilo kojem zahtjevu od 1 do 4 i 6, prije sijanja i/ili nakon pred-klijanja, s AHAS-inhibirajućim imidazolinon-herbicidom, ili s mješavinom imidazolinon-herbicida, sulfonilurea-herbicida i/ili triazolopirimidin-herbicida.
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| PCT/US2006/025534 WO2007005581A2 (en) | 2005-07-01 | 2006-06-29 | Herbicide-resistant sunflower plants, polynucleotides encoding herbicide=resistant acetohydroxyacid synthase large subunit proteins, and methods of use |
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| HRP20120294TT HRP20120294T1 (hr) | 2005-07-01 | 2012-04-03 | Suncokretove biljke otporne na herbicide, za polinukleotide koji kodiraju velike pojedinačne proteine otporne na herbicide sintezom acetohidroksilne kiseline, te postupci uporabe |
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