HRP20171955T1 - Suncokreti otporni na herbicide s više alela ahasl1 otpornih na herbicide i postupci njihove upotrebe - Google Patents
Suncokreti otporni na herbicide s više alela ahasl1 otpornih na herbicide i postupci njihove upotrebe Download PDFInfo
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- HRP20171955T1 HRP20171955T1 HRP20171955TT HRP20171955T HRP20171955T1 HR P20171955 T1 HRP20171955 T1 HR P20171955T1 HR P20171955T T HRP20171955T T HR P20171955TT HR P20171955 T HRP20171955 T HR P20171955T HR P20171955 T1 HRP20171955 T1 HR P20171955T1
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- herbicide
- ahasl1
- methyl
- sunflower
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- 230000002363 herbicidal effect Effects 0.000 title claims 44
- 239000004009 herbicide Substances 0.000 title claims 44
- 241000208818 Helianthus Species 0.000 title claims 21
- 238000000034 method Methods 0.000 title claims 19
- 108700028369 Alleles Proteins 0.000 title claims 15
- 235000003222 Helianthus annuus Nutrition 0.000 claims 20
- 239000002773 nucleotide Substances 0.000 claims 17
- 125000003729 nucleotide group Chemical group 0.000 claims 17
- 108090000623 proteins and genes Proteins 0.000 claims 15
- 101710103719 Acetolactate synthase large subunit Proteins 0.000 claims 10
- 101710182467 Acetolactate synthase large subunit IlvB1 Proteins 0.000 claims 10
- 101710171176 Acetolactate synthase large subunit IlvG Proteins 0.000 claims 10
- 101710196435 Probable acetolactate synthase large subunit Proteins 0.000 claims 10
- 125000003275 alpha amino acid group Chemical group 0.000 claims 10
- 235000001014 amino acid Nutrition 0.000 claims 10
- 238000006467 substitution reaction Methods 0.000 claims 10
- 239000000203 mixture Substances 0.000 claims 9
- CAAMSDWKXXPUJR-UHFFFAOYSA-N 3,5-dihydro-4H-imidazol-4-one Chemical compound O=C1CNC=N1 CAAMSDWKXXPUJR-UHFFFAOYSA-N 0.000 claims 8
- 235000018102 proteins Nutrition 0.000 claims 8
- 102000004169 proteins and genes Human genes 0.000 claims 8
- 235000004279 alanine Nutrition 0.000 claims 7
- 229940100389 Sulfonylurea Drugs 0.000 claims 5
- YROXIXLRRCOBKF-UHFFFAOYSA-N sulfonylurea Chemical class OC(=N)N=S(=O)=O YROXIXLRRCOBKF-UHFFFAOYSA-N 0.000 claims 5
- PRZRAMLXTKZUHF-UHFFFAOYSA-N 5-oxo-n-sulfonyl-4h-triazole-1-carboxamide Chemical compound O=S(=O)=NC(=O)N1N=NCC1=O PRZRAMLXTKZUHF-UHFFFAOYSA-N 0.000 claims 4
- 241000196324 Embryophyta Species 0.000 claims 4
- QNAYBMKLOCPYGJ-REOHCLBHSA-N L-alanine Chemical compound C[C@H](N)C(O)=O QNAYBMKLOCPYGJ-REOHCLBHSA-N 0.000 claims 4
- AYFVYJQAPQTCCC-UHFFFAOYSA-N Threonine Natural products CC(O)C(N)C(O)=O AYFVYJQAPQTCCC-UHFFFAOYSA-N 0.000 claims 4
- 239000004473 Threonine Substances 0.000 claims 4
- YWBFPKPWMSWWEA-UHFFFAOYSA-O triazolopyrimidine Chemical compound BrC1=CC=CC(C=2N=C3N=CN[N+]3=C(NCC=3C=CN=CC=3)C=2)=C1 YWBFPKPWMSWWEA-UHFFFAOYSA-O 0.000 claims 4
- CLQMBPJKHLGMQK-UHFFFAOYSA-N 2-(4-isopropyl-4-methyl-5-oxo-4,5-dihydro-1H-imidazol-2-yl)nicotinic acid Chemical compound N1C(=O)C(C(C)C)(C)N=C1C1=NC=CC=C1C(O)=O CLQMBPJKHLGMQK-UHFFFAOYSA-N 0.000 claims 3
- ROHFNLRQFUQHCH-UHFFFAOYSA-N Leucine Natural products CC(C)CC(N)C(O)=O ROHFNLRQFUQHCH-UHFFFAOYSA-N 0.000 claims 3
- ONIBWKKTOPOVIA-UHFFFAOYSA-N Proline Natural products OC(=O)C1CCCN1 ONIBWKKTOPOVIA-UHFFFAOYSA-N 0.000 claims 3
- KZSNJWFQEVHDMF-UHFFFAOYSA-N Valine Natural products CC(C)C(N)C(O)=O KZSNJWFQEVHDMF-UHFFFAOYSA-N 0.000 claims 3
- 150000001294 alanine derivatives Chemical class 0.000 claims 3
- 230000000295 complement effect Effects 0.000 claims 3
- 230000002401 inhibitory effect Effects 0.000 claims 3
- 235000020238 sunflower seed Nutrition 0.000 claims 3
- 239000004474 valine Substances 0.000 claims 3
- NUPJIGQFXCQJBK-UHFFFAOYSA-N 2-(4-isopropyl-4-methyl-5-oxo-4,5-dihydro-1H-imidazol-2-yl)-5-(methoxymethyl)nicotinic acid Chemical compound OC(=O)C1=CC(COC)=CN=C1C1=NC(C)(C(C)C)C(=O)N1 NUPJIGQFXCQJBK-UHFFFAOYSA-N 0.000 claims 2
- PVSGXWMWNRGTKE-UHFFFAOYSA-N 5-methyl-2-[4-methyl-5-oxo-4-(propan-2-yl)-4,5-dihydro-1H-imidazol-2-yl]pyridine-3-carboxylic acid Chemical compound N1C(=O)C(C(C)C)(C)N=C1C1=NC=C(C)C=C1C(O)=O PVSGXWMWNRGTKE-UHFFFAOYSA-N 0.000 claims 2
- XVOKUMIPKHGGTN-UHFFFAOYSA-N Imazethapyr Chemical compound OC(=O)C1=CC(CC)=CN=C1C1=NC(C)(C(C)C)C(=O)N1 XVOKUMIPKHGGTN-UHFFFAOYSA-N 0.000 claims 2
- 241001613787 Orobanche grenieri Species 0.000 claims 2
- 238000012408 PCR amplification Methods 0.000 claims 2
- 125000001909 leucine group Chemical group [H]N(*)C(C(*)=O)C([H])([H])C(C([H])([H])[H])C([H])([H])[H] 0.000 claims 2
- FFCCBBNQPIMUJI-UHFFFAOYSA-N methyl 2-(4-isopropyl-4-methyl-5-oxo-2-imidazolin-2-yl)-para-toluate Chemical compound COC(=O)C1=CC=C(C)C=C1C1=NC(C)(C(C)C)C(=O)N1 FFCCBBNQPIMUJI-UHFFFAOYSA-N 0.000 claims 2
- TYIHVCIQMMTVHE-UHFFFAOYSA-N methyl 2-(4-isopropyl-4-methyl-5-oxo-4,5-dihydro-1H-imidazol-2-yl)-5-methylbenzoate Chemical compound COC(=O)C1=CC(C)=CC=C1C1=NC(C)(C(C)C)C(=O)N1 TYIHVCIQMMTVHE-UHFFFAOYSA-N 0.000 claims 2
- 238000003752 polymerase chain reaction Methods 0.000 claims 2
- 125000000341 threoninyl group Chemical group [H]OC([H])(C([H])([H])[H])C([H])(N([H])[H])C(*)=O 0.000 claims 2
- 239000001226 triphosphate Substances 0.000 claims 2
- 235000011178 triphosphate Nutrition 0.000 claims 2
- 125000002264 triphosphate group Chemical class [H]OP(=O)(O[H])OP(=O)(O[H])OP(=O)(O[H])O* 0.000 claims 2
- 125000002987 valine group Chemical group [H]N([H])C([H])(C(*)=O)C([H])(C([H])([H])[H])C([H])([H])[H] 0.000 claims 2
- WRIDQFICGBMAFQ-UHFFFAOYSA-N (E)-8-Octadecenoic acid Natural products CCCCCCCCCC=CCCCCCCC(O)=O WRIDQFICGBMAFQ-UHFFFAOYSA-N 0.000 claims 1
- LQJBNNIYVWPHFW-UHFFFAOYSA-N 20:1omega9c fatty acid Natural products CCCCCCCCCCC=CCCCCCCCC(O)=O LQJBNNIYVWPHFW-UHFFFAOYSA-N 0.000 claims 1
- QSBYPNXLFMSGKH-UHFFFAOYSA-N 9-Heptadecensaeure Natural products CCCCCCCC=CCCCCCCCC(O)=O QSBYPNXLFMSGKH-UHFFFAOYSA-N 0.000 claims 1
- 239000003666 Amidosulfuron Substances 0.000 claims 1
- CTTHWASMBLQOFR-UHFFFAOYSA-N Amidosulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)N(C)S(C)(=O)=O)=N1 CTTHWASMBLQOFR-UHFFFAOYSA-N 0.000 claims 1
- 239000005472 Bensulfuron methyl Substances 0.000 claims 1
- 239000005496 Chlorsulfuron Substances 0.000 claims 1
- WMLPCIHUFDKWJU-UHFFFAOYSA-N Cinosulfuron Chemical compound COCCOC1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(OC)=NC(OC)=N1 WMLPCIHUFDKWJU-UHFFFAOYSA-N 0.000 claims 1
- 239000005514 Flazasulfuron Substances 0.000 claims 1
- HWATZEJQIXKWQS-UHFFFAOYSA-N Flazasulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)C=2C(=CC=CN=2)C(F)(F)F)=N1 HWATZEJQIXKWQS-UHFFFAOYSA-N 0.000 claims 1
- LXKOADMMGWXPJQ-UHFFFAOYSA-N Halosulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)C=2N(N=C(Cl)C=2C(O)=O)C)=N1 LXKOADMMGWXPJQ-UHFFFAOYSA-N 0.000 claims 1
- 239000005567 Imazosulfuron Substances 0.000 claims 1
- NAGRVUXEKKZNHT-UHFFFAOYSA-N Imazosulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)C=2N3C=CC=CC3=NC=2Cl)=N1 NAGRVUXEKKZNHT-UHFFFAOYSA-N 0.000 claims 1
- ROHFNLRQFUQHCH-YFKPBYRVSA-N L-leucine Chemical compound CC(C)C[C@H](N)C(O)=O ROHFNLRQFUQHCH-YFKPBYRVSA-N 0.000 claims 1
- KZSNJWFQEVHDMF-BYPYZUCNSA-N L-valine Chemical compound CC(C)[C@H](N)C(O)=O KZSNJWFQEVHDMF-BYPYZUCNSA-N 0.000 claims 1
- 239000005584 Metsulfuron-methyl Substances 0.000 claims 1
- 239000005586 Nicosulfuron Substances 0.000 claims 1
- 239000005642 Oleic acid Substances 0.000 claims 1
- ZQPPMHVWECSIRJ-UHFFFAOYSA-N Oleic acid Natural products CCCCCCCCC=CCCCCCCCC(O)=O ZQPPMHVWECSIRJ-UHFFFAOYSA-N 0.000 claims 1
- 235000009540 Orobanche cernua Nutrition 0.000 claims 1
- 235000001487 Orobanche cernua var cumana Nutrition 0.000 claims 1
- BGNQYGRXEXDAIQ-UHFFFAOYSA-N Pyrazosulfuron-ethyl Chemical group C1=NN(C)C(S(=O)(=O)NC(=O)NC=2N=C(OC)C=C(OC)N=2)=C1C(=O)OCC BGNQYGRXEXDAIQ-UHFFFAOYSA-N 0.000 claims 1
- 239000005616 Rimsulfuron Substances 0.000 claims 1
- 239000005623 Thifensulfuron-methyl Substances 0.000 claims 1
- 230000003321 amplification Effects 0.000 claims 1
- XMQFTWRPUQYINF-UHFFFAOYSA-N bensulfuron-methyl Chemical group COC(=O)C1=CC=CC=C1CS(=O)(=O)NC(=O)NC1=NC(OC)=CC(OC)=N1 XMQFTWRPUQYINF-UHFFFAOYSA-N 0.000 claims 1
- NSWAMPCUPHPTTC-UHFFFAOYSA-N chlorimuron-ethyl Chemical group CCOC(=O)C1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(Cl)=CC(OC)=N1 NSWAMPCUPHPTTC-UHFFFAOYSA-N 0.000 claims 1
- VJYIFXVZLXQVHO-UHFFFAOYSA-N chlorsulfuron Chemical compound COC1=NC(C)=NC(NC(=O)NS(=O)(=O)C=2C(=CC=CC=2)Cl)=N1 VJYIFXVZLXQVHO-UHFFFAOYSA-N 0.000 claims 1
- 238000003205 genotyping method Methods 0.000 claims 1
- 230000035784 germination Effects 0.000 claims 1
- QXJSBBXBKPUZAA-UHFFFAOYSA-N isooleic acid Natural products CCCCCCCC=CCCCCCCCCC(O)=O QXJSBBXBKPUZAA-UHFFFAOYSA-N 0.000 claims 1
- ZTYVMAQSHCZXLF-UHFFFAOYSA-N methyl 2-[[4,6-bis(difluoromethoxy)pyrimidin-2-yl]carbamoylsulfamoyl]benzoate Chemical group COC(=O)C1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(OC(F)F)=CC(OC(F)F)=N1 ZTYVMAQSHCZXLF-UHFFFAOYSA-N 0.000 claims 1
- RSMUVYRMZCOLBH-UHFFFAOYSA-N metsulfuron methyl Chemical group COC(=O)C1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(C)=NC(OC)=N1 RSMUVYRMZCOLBH-UHFFFAOYSA-N 0.000 claims 1
- RTCOGUMHFFWOJV-UHFFFAOYSA-N nicosulfuron Chemical compound COC1=CC(OC)=NC(NC(=O)NS(=O)(=O)C=2C(=CC=CN=2)C(=O)N(C)C)=N1 RTCOGUMHFFWOJV-UHFFFAOYSA-N 0.000 claims 1
- 238000003199 nucleic acid amplification method Methods 0.000 claims 1
- ZQPPMHVWECSIRJ-KTKRTIGZSA-N oleic acid Chemical compound CCCCCCCC\C=C/CCCCCCCC(O)=O ZQPPMHVWECSIRJ-KTKRTIGZSA-N 0.000 claims 1
- 125000001500 prolyl group Chemical class [H]N1C([H])(C(=O)[*])C([H])([H])C([H])([H])C1([H])[H] 0.000 claims 1
- DJXNJVFEFSWHLY-UHFFFAOYSA-N quinoline-3-carboxylic acid Chemical compound C1=CC=CC2=CC(C(=O)O)=CN=C21 DJXNJVFEFSWHLY-UHFFFAOYSA-N 0.000 claims 1
- MEFOUWRMVYJCQC-UHFFFAOYSA-N rimsulfuron Chemical compound CCS(=O)(=O)C1=CC=CN=C1S(=O)(=O)NC(=O)NC1=NC(OC)=CC(OC)=N1 MEFOUWRMVYJCQC-UHFFFAOYSA-N 0.000 claims 1
- 238000009331 sowing Methods 0.000 claims 1
- ZDXMLEQEMNLCQG-UHFFFAOYSA-N sulfometuron methyl Chemical group COC(=O)C1=CC=CC=C1S(=O)(=O)NC(=O)NC1=NC(C)=CC(C)=N1 ZDXMLEQEMNLCQG-UHFFFAOYSA-N 0.000 claims 1
- AHTPATJNIAFOLR-UHFFFAOYSA-N thifensulfuron-methyl Chemical group S1C=CC(S(=O)(=O)NC(=O)NC=2N=C(OC)N=C(C)N=2)=C1C(=O)OC AHTPATJNIAFOLR-UHFFFAOYSA-N 0.000 claims 1
- XOPFESVZMSQIKC-UHFFFAOYSA-N triasulfuron Chemical compound COC1=NC(C)=NC(NC(=O)NS(=O)(=O)C=2C(=CC=CC=2)OCCCl)=N1 XOPFESVZMSQIKC-UHFFFAOYSA-N 0.000 claims 1
- YMXOXAPKZDWXLY-QWRGUYRKSA-N tribenuron methyl Chemical group COC(=O)[C@H]1CCCC[C@@H]1S(=O)(=O)NC(=O)N(C)C1=NC(C)=NC(OC)=N1 YMXOXAPKZDWXLY-QWRGUYRKSA-N 0.000 claims 1
- IMEVJVISCHQJRM-UHFFFAOYSA-N triflusulfuron-methyl Chemical group COC(=O)C1=CC=CC(C)=C1S(=O)(=O)NC(=O)NC1=NC(OCC(F)(F)F)=NC(N(C)C)=N1 IMEVJVISCHQJRM-UHFFFAOYSA-N 0.000 claims 1
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- A—HUMAN NECESSITIES
- A01—AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
- A01H—NEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
- A01H5/00—Angiosperms, i.e. flowering plants, characterised by their plant parts; Angiosperms characterised otherwise than by their botanic taxonomy
- A01H5/02—Flowers
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- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8241—Phenotypically and genetically modified plants via recombinant DNA technology
- C12N15/8261—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield
- C12N15/8271—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield for stress resistance, e.g. heavy metal resistance
- C12N15/8274—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield for stress resistance, e.g. heavy metal resistance for herbicide resistance
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- A—HUMAN NECESSITIES
- A01—AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
- A01H—NEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
- A01H1/00—Processes for modifying genotypes ; Plants characterised by associated natural traits
- A01H1/12—Processes for modifying agronomic input traits, e.g. crop yield
- A01H1/122—Processes for modifying agronomic input traits, e.g. crop yield for stress resistance, e.g. heavy metal resistance
- A01H1/123—Processes for modifying agronomic input traits, e.g. crop yield for stress resistance, e.g. heavy metal resistance for herbicide resistance
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- A—HUMAN NECESSITIES
- A01—AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
- A01H—NEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
- A01H6/00—Angiosperms, i.e. flowering plants, characterised by their botanic taxonomy
- A01H6/14—Asteraceae or Compositae, e.g. safflower, sunflower, artichoke or lettuce
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- A—HUMAN NECESSITIES
- A01—AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
- A01N—PRESERVATION OF BODIES OF HUMANS OR ANIMALS OR PLANTS OR PARTS THEREOF; BIOCIDES, e.g. AS DISINFECTANTS, AS PESTICIDES OR AS HERBICIDES; PEST REPELLANTS OR ATTRACTANTS; PLANT GROWTH REGULATORS
- A01N43/00—Biocides, pest repellants or attractants, or plant growth regulators containing heterocyclic compounds
- A01N43/48—Biocides, pest repellants or attractants, or plant growth regulators containing heterocyclic compounds having rings with two nitrogen atoms as the only ring hetero atoms
- A01N43/50—1,3-Diazoles; Hydrogenated 1,3-diazoles
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- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8241—Phenotypically and genetically modified plants via recombinant DNA technology
- C12N15/8261—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield
- C12N15/8271—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield for stress resistance, e.g. heavy metal resistance
- C12N15/8274—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield for stress resistance, e.g. heavy metal resistance for herbicide resistance
- C12N15/8278—Sulfonylurea
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- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
- C12N9/88—Lyases (4.)
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Claims (22)
1. Suncokret otporan na herbicide, naznačen time što sadrži dva različita alela otporna na herbicide gena za veliku podjedinicu 1 suncokretske sintaze acetohidroksikiselina (AHASL1), gdje prvi alel kodira prvi protein AHASL1 otporan na herbicide, koji sadrži zamjenu alanina treoninom na aminokiselinskom položaju koji odgovara položaju 7 u SEQ ID NO:20, a drugi alel kodira drugi protein AHASL1 otporan na herbicide, koji sadrži zamjenu alanina valinom na aminokiselinskom položaju koji odgovara položaju 90 u SEQ ID NO:20 ili zamjenu prolina leucinom na aminokiselinskom položaju koji odgovara položaju 82 u SEQ ID NO:20; gdje je navedeni suncokret otporan na herbicide otporan na najmanje jedan herbicid koji inhibira AHAS.
2. Suncokret u skladu s patentnim zahtjevom 1, naznačen time što je navedeni herbicid koji inhibira AHAS imidazolinonski herbicid, sulfonilurejski herbicid, triazolopirimidinski herbicid, sulfonilamino-karboniltriazolinonski herbicid, ili njihove smjese.
3. Suncokret u skladu s patentnim zahtjevom 1, naznačen time što je navedeni suncokret u obliku sjemena.
4. Suncokret u skladu s patentnim zahtjevom 1, naznačen time što navedeni suncokret proizvodi sjemenke koje sadrže ulje koje se može ekstrahirati iz sjemenki koje sadrži najmanje 85% oleinske kiseline.
5. Postupak uzgoja suncokreta prilikom kontrole korova u blizini navedenog suncokreta, gdje se navedeni postupak sastoji u primjeni djelotvorne količina herbicide koji inhibira AHAS na korovima i na suncokretu, naznačen time što se navedeni suncokret bira iz skupine koju čine:
(a) suncokret koji sadrži prvi alel gena AHASL1 i drugi alel navedenog gena AHASL1, gdje navedeni prvi alel kodira prvi protein AHASL1 otporan na herbicide, koji sadrži zamjenu alanina treoninom na aminokiselinskom položaju koji odgovara položaju 7 u SEQ ID NO:20, a navedeni drugi alel kodira drugi protein AHASL1 otporan na herbicide, koji sadrži zamjenu alanina valinom na aminokiselinskom položaju koji odgovara položaju 90 u SEQ ID NO:20; i
(b) suncokret koji sadrži prvi alel gena AHASL1 i drugi alel navedenog gena AHASL1, gdje navedeni prvi alel kodira prvi protein AHASL1 otporan na herbicide, koji sadrži zamjenu alanina treoninom na aminokiselinskom položaju koji odgovara položaju 7 u SEQ ID NO:20, a navedeni drugi alel kodira drugi protein AHASL1 otporan na herbicide, koji sadrži zamjena prolina leucinom na aminokiselinskom položaju koji odgovara položaju 82 u SEQ ID NO:20.
6. Postupak u skladu s patentnim zahtjevom 5, naznačen time što navedeni herbicid koji inhibira AHAS sadrži imidazolinonski herbicid, sulfonilurejski herbicid, triazolopirimidinski herbicid, sulfonilamino-karboniltriazolinonski herbicid, ili njihova smjesa.
7. Postupak u skladu s patentnim zahtjevom 6, naznačen time što navedeni imidazolinonski herbicid sadrži: 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)nikotinsku kiselinu, 2-(4-izopropil)-4-metil-5-okso-2-imidazolin-2-il)kinolin-3-karboksilnu kiselinu, 5-etil-2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)nikotinsku kiselinu, 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-5-(metoksimetil)nikotinsku kiselinu, 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-5-metilnikotinsku kiselinu, smjese metil-6-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-m-toluata i metil-2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-p-toluata, ili njihove smjese.
8. Postupak u skladu s patentnim zahtjevom 6, naznačen time što navedeni sulfonilurejski herbicid sadrži: klorsulfuron, metsulfuron-metil, sulfometuron-metil, klorimuron-etil, tifensulfuron-metil, tribenuron-metil, bensulfuron-metil, nicosulfuron, etametsulfuron-metil, rimsulfuron, triflusulfuron-metil, triasulfuron, primisulfuron-metil, cinosulfuron, amidosulfuron, flazasulfuron, imazosulfuron, pirazosulfuron-etil, halosulfuron, ili njihove smjese.
9. Sjeme suncokreta u skladu s patentnim zahtjevom 3, naznačeno time što se navedeno sjeme tretira herbicidom koji inhibira AHAS.
10. Sjeme u skladu s patentnim zahtjevom 9, naznačeno time što navedeni herbicid koji inhibira AHAS sadrži imidazolinonski herbicid, sulfonilurejski herbicid, triazolopirimidinski herbicid, sulfonilamino-karboniltriazolinonski herbicid, ili njihovu smjesu.
11. Postupak uzgoja suncokreta otpornog na herbicide u skladu s patentnim zahtjevom 1 prilikom borbe protiv nepoželjnog raslinja, naznačen time što se sastoji u stavljanju u kontakt sjemena suncokreta prije sjetve i/ili nakon predklijanja djelotvornom količinom herbicida koji inhibira AHAS, gdje navedeno sjeme suncokreta sadrži dva različita alela otporna na herbicide gena za veliku podjedinicu 1 suncokretske sintaze acetohidroksikiselina (AHASL1), gdje prvi alel kodira prvi protein AHASL1 otporan na herbicide, koji sadrži zamjenu alanina treoninom na aminokiselinskom položaju koji odgovara položaju 7 u SEQ ID NO:20, a drugi alel kodira drugi protein AHASL1 otporan na herbicide, koji sadrži zamjenu alanina valinom na aminokiselinskom položaju koji odgovara položaju 90 u SEQ ID NO:20 ili zamjenu prolina leucinom na aminokiselinskom položaju koji odgovara položaju 82 u SEQ ID NO:20.
12. Postupak u skladu s patentnim zahtjevom 11, naznačen time što navedeni herbicid koji inhibira AHAS sadrži imidazolinonski herbicid, sulfonilurejski herbicid, triazolopirimidinski herbicid, sulfonilamino-karboniltriazolinonski herbicid, ili njihovu smjesu.
13. Postupak u skladu s patentnim zahtjevom 5, naznačen time što navedeni korovi uključuju volovod.
14. Postupak u skladu s patentnim zahtjevom 13, naznačen time što se navedeni volovod bira iz skupine koju čine: Orobanche cumana i Orobanche cernua.
15. Postupak u skladu s patentnim zahtjevom 13, naznačen time što navedeni imidazolinonski herbicid sadrži: 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)nikotinsku kiselinu, 2-(4-izopropil)-4-metil-5-okso-2-imidazolin-2-il)-3-kinolinkarboksilnu kiselinu, 5-etil-2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)nikotinsku kiselinu, 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-5-(metoksimetil)nikotinsku kiselinu, 2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-5-metilnikotinsku kiselinu, smjese metil-6-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-m-toluata i metil-2-(4-izopropil-4-metil-5-okso-2-imidazolin-2-il)-p-toluata, ili njihove smjese.
16. Postupak u skladu s patentnim zahtjevom 13, naznačen time što je navedeni imidazolinonski herbicid imazapir.
17. Postupak u skladu s patentnim zahtjevom 13, naznačen time što se navedeni imidazolinonski herbicid primjenjuje u stadiju rasta R1.
18. Postupak genotipiziranja suncokreta u skladu s bilo kojim od patentnih zahtjeva 1-4, ili njegovog potomstva, ili sjemena u skladu s bilo kojim od patentnih zahtjeva 9-10, što sadrži suncokretski AHASL1, naznačen time što se navedeni postupak sastoji u koracima:
(a) dobivanja genomske DNA iz navedenog suncokreta ili sjemena;
(b) upotrebe navedene DNA kao uzorka za prvu amplifikaciju lančanom reakcijom polimeraze (PCR), koja uključuje navedenu DNA, polimerazu, deosiribonukleotid-trifosfate, početnicu prema naprijed za AHASL1 i početnicu prema natrag za divlji tip AHASL1, gdje se navedena početnica prema natrag za divlji tip AHASL1 sparuje s nukleotidnim slijedom koji sadrži nukleotidni slijed iznijet u SEQ ID NO:13, gdje je nukleotid koji je na 3′ kraju nukleotid navedene početnice prema natrag za divlji tip AHASL1 koji je komplement nukleotida koji je na položaju 1 u nukleotidnom slijedu iznijetom u SEQ ID NO:13;
(c) upotreba navedene DNA kao uzorak za drugu PCR amplifikaciju koja uključuje navedenu DNA, polimerazu, deosiribonukleotid-trifosfate, navedenu početnicu prema naprijed za AHASL1 i početnicu prema natrag za mutantni AHASL1, gdje se navedena početnica prema natrag za mutantni AHASL1 sparuje s nukleotidnim slijedom koji sadrži nukleotidni slijed iznijet u SEQ ID NO:14, gdje je nukleotid koji je na 3′ kraju nukleotid navedene početnice prema natrag za mutantni AHASL1 koji je komplement nukleotida koji je na položaju 1 u nukleotidnom slijedu iznijetom u SEQ ID NO:14; i
(d) detektiranja produkata navedene prve i navedene druge PCR amplifikacije;
gdje navedena početnica prema naprijed za AHASL1 sadrži nukleotidni slijed koji odgovara području suncokretskog gena AHASL1 koje je 5′ u odnosu na područje (ACC)n.
19. Postupak u skladu s patentnim zahtjevom 18, naznačen time što navedena početnica prema natrag za divlji tip AHASL1 sadrži nukleotidni slijed iznijet u SEQ ID NO:4.
20. Postupak u skladu s patentnim zahtjevom 18, naznačen time što navedena početnica prema natrag za mutantni AHASL1 sadrži nukleotidni slijed iznijet u SEQ ID NO:5.
21. Postupak u skladu s patentnim zahtjevom 18, naznačen time što se navedena početnica prema naprijed za AHASL1 sparuje s nukleotidnim slijedom koji sadrži komplement nukleotidnog slijeda iznijetog u SEQ ID NO:12.
22. Postupak u skladu s patentnim zahtjevom 18, naznačen time što navedena početnica prema naprijed za AHASL1 sadrži nukleotidni slijed iznijet u SEQ ID NO:3.
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| US91004107P | 2007-04-04 | 2007-04-04 | |
| US2973708P | 2008-02-19 | 2008-02-19 | |
| EP08744931.0A EP2134839B8 (en) | 2007-04-04 | 2008-04-02 | Herbicide-resistant sunflower plants with multiple herbicide resistant alleles of ahasl1 and methods of use |
| PCT/US2008/059125 WO2008124431A1 (en) | 2007-04-04 | 2008-04-02 | Herbicide-resistant sunflower plants with multiple herbicide resistant alleles of ahasl1 and methods of use |
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| ES2692594T1 (es) * | 2005-03-02 | 2018-12-04 | Instituto Nacional De Tecnologia Agropecuaria | Plantas de arroz resistentes a herbicidas, polinucleótidos que codifican proteínas de la subunidad grande de la acetohidroxiácido sintasa resistentes a herbicidas y métodos para su uso |
| UA115430C2 (uk) * | 2011-09-13 | 2017-11-10 | Басф Агрокемікал Продактс Б.В. | Спосіб боротьби з паразитними бур'янами за допомогою сумішей, що включають гербіцидні інгібітори ацетолактатсинтази й регулятори росту рослин |
| WO2013081987A1 (en) * | 2011-11-29 | 2013-06-06 | Agrigenetics, Inc. | High throughput single nucleotide polymorphism assay |
| BR112014013486A8 (pt) * | 2011-12-05 | 2017-06-27 | Basf Agrochemical Products Bv | método de controle de vegetação indesejável e uso de composições |
| MX366944B (es) * | 2012-04-05 | 2019-07-31 | Advanta Int Bv | Plantas de sorgo que tienen un polinucleotido mutado que codifica una subunidad mayor de la proteina sintasa acetohidroxiacida mutada y tiene resistencia incrementada a herbicidas. |
| US20150148235A1 (en) * | 2013-11-22 | 2015-05-28 | Nidera S.A. | Herbicide-resistant sunflower plants with multiple herbicide resistant alleles of ahasl1 and methods of use |
| US20150148234A1 (en) * | 2013-11-22 | 2015-05-28 | Nidera S.A. | Herbicide-resistant sunflower plants with multiple herbicide resistant alleles of ahasl1 and methods of use |
| JP2017118878A (ja) * | 2017-01-16 | 2017-07-06 | アドヴァンタ・ホールディングス・ベスローテン・フェンノートシャップ | 突然変異アセトヒドロキシ酸合成酵素タンパク質のラージサブユニットをコードする突然変異ポリヌクレオチドを有し、除草剤耐性が増大したソルガム植物 |
| WO2018187543A1 (en) * | 2017-04-06 | 2018-10-11 | Dow Agrosciences Llc | Molecular markers associated with orobanche resistance in sunflower |
| CN111690625B (zh) * | 2017-07-13 | 2023-04-11 | 江苏省农业科学院 | 具有除草剂抗性的乙酰乳酸合酶突变蛋白及其应用 |
| WO2019165411A1 (en) * | 2018-02-26 | 2019-08-29 | Just Biotherapeutics, Inc. | Determining impact on properties of proteins based on amino acid sequence modifications |
| CN108739354A (zh) * | 2018-05-12 | 2018-11-06 | 新疆农业科学院经济作物研究所 | 抗g型列当向日葵恢复系1808r的选育方法 |
| CN109315235A (zh) * | 2018-09-30 | 2019-02-12 | 新疆生产建设兵团第十师农业科学研究所 | 一种感列当向日葵品种的抗列当改良方法 |
| CA3130265A1 (en) | 2019-02-19 | 2020-08-27 | Gowan Company, L.L.C. | Stable liquid compositions and methods of using the same |
| JP2019141045A (ja) * | 2019-03-04 | 2019-08-29 | アドヴァンタ・ホールディングス・ベスローテン・フェンノートシャップ | 突然変異アセトヒドロキシ酸合成酵素タンパク質のラージサブユニットをコードする突然変異ポリヌクレオチドを有し、除草剤耐性が増大したソルガム植物 |
| AR127152A1 (es) | 2021-09-27 | 2023-12-20 | BASF Agricultural Solutions Seed US LLC | Plantas de girasol no transgénicas que tienen tolerancia aumentada a herbicidas |
| CN114606333A (zh) * | 2022-03-16 | 2022-06-10 | 华智生物技术有限公司 | 用于鉴定耐磺酰脲类除草剂向日葵的引物组、试剂盒及其应用 |
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| US5545822A (en) | 1992-08-21 | 1996-08-13 | Board Of Supervisors Of Louisiana State University And Agricultural And Mechanical College | Herbicide resistant rice |
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| US7432082B2 (en) * | 2004-03-22 | 2008-10-07 | Basf Ag | Methods and compositions for analyzing AHASL genes |
| RU2415566C2 (ru) * | 2004-07-30 | 2011-04-10 | Басф Агрокемикал Продактс Б.В. | Устойчивые к гербицидам растения подсолнечника, полинуклеотиды, кодирующие устойчивые к гербицидам белки большой субъединицы ацетогидроксиацидсинтазы, и способы применения |
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| CL2012003055A1 (es) | 2013-02-15 |
| CA2682554A1 (en) | 2008-10-16 |
| US20110277051A1 (en) | 2011-11-10 |
| CL2008000975A1 (es) | 2009-03-27 |
| EA019952B1 (ru) | 2014-07-30 |
| SI2134839T1 (en) | 2018-05-31 |
| CN102808017A (zh) | 2012-12-05 |
| JP5715414B2 (ja) | 2015-05-07 |
| BRPI0809473A2 (pt) | 2014-09-09 |
| JP6117761B2 (ja) | 2017-04-19 |
| EP2134839B1 (en) | 2017-12-06 |
| AU2008237436B2 (en) | 2014-01-23 |
| WO2008124431A1 (en) | 2008-10-16 |
| CA2682554C (en) | 2021-08-10 |
| EA200901268A1 (ru) | 2010-06-30 |
| JP2010523122A (ja) | 2010-07-15 |
| AU2008237436A1 (en) | 2008-10-16 |
| ES2659991T3 (es) | 2018-03-20 |
| PT2134839T (pt) | 2018-01-15 |
| EP2134839A1 (en) | 2009-12-23 |
| AR065953A1 (es) | 2009-07-15 |
| US20160222405A1 (en) | 2016-08-04 |
| EA201270282A1 (ru) | 2012-07-30 |
| ZA200907701B (en) | 2013-02-27 |
| CN102808017B (zh) | 2016-01-13 |
| JP2015057060A (ja) | 2015-03-26 |
| US20160222404A1 (en) | 2016-08-04 |
| CN101878300A (zh) | 2010-11-03 |
| EP2134839B8 (en) | 2018-08-15 |
| EA023348B1 (ru) | 2016-05-31 |
| EA019952B9 (ru) | 2015-01-30 |
| CN101878300B (zh) | 2013-09-11 |
| UA102223C2 (ru) | 2013-06-25 |
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