JP6591993B2 - グルコースシロップ生産用組成物 - Google Patents

グルコースシロップ生産用組成物 Download PDF

Info

Publication number
JP6591993B2
JP6591993B2 JP2016550206A JP2016550206A JP6591993B2 JP 6591993 B2 JP6591993 B2 JP 6591993B2 JP 2016550206 A JP2016550206 A JP 2016550206A JP 2016550206 A JP2016550206 A JP 2016550206A JP 6591993 B2 JP6591993 B2 JP 6591993B2
Authority
JP
Japan
Prior art keywords
seq
amylase
pullulanase
glucoamylase
mature polypeptide
Prior art date
Legal status (The legal status is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the status listed.)
Active
Application number
JP2016550206A
Other languages
English (en)
Japanese (ja)
Other versions
JP2017505129A (ja
JP2017505129A5 (enExample
Inventor
志朗 福山
志朗 福山
圭一 綾部
圭一 綾部
Original Assignee
ノボザイムス アクティーゼルスカブ
ノボザイムス アクティーゼルスカブ
Priority date (The priority date is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the date listed.)
Filing date
Publication date
Application filed by ノボザイムス アクティーゼルスカブ, ノボザイムス アクティーゼルスカブ filed Critical ノボザイムス アクティーゼルスカブ
Publication of JP2017505129A publication Critical patent/JP2017505129A/ja
Publication of JP2017505129A5 publication Critical patent/JP2017505129A5/ja
Application granted granted Critical
Publication of JP6591993B2 publication Critical patent/JP6591993B2/ja
Active legal-status Critical Current
Anticipated expiration legal-status Critical

Links

Classifications

    • AHUMAN NECESSITIES
    • A23FOODS OR FOODSTUFFS; TREATMENT THEREOF, NOT COVERED BY OTHER CLASSES
    • A23LFOODS, FOODSTUFFS OR NON-ALCOHOLIC BEVERAGES, NOT OTHERWISE PROVIDED FOR; PREPARATION OR TREATMENT THEREOF
    • A23L29/00Foods or foodstuffs containing additives; Preparation or treatment thereof
    • A23L29/30Foods or foodstuffs containing additives; Preparation or treatment thereof containing carbohydrate syrups; containing sugars; containing sugar alcohols, e.g. xylitol; containing starch hydrolysates, e.g. dextrin
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12PFERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P19/00Preparation of compounds containing saccharide radicals
    • C12P19/14Preparation of compounds containing saccharide radicals produced by the action of a carbohydrase (EC 3.2.x), e.g. by alpha-amylase, e.g. by cellulase, hemicellulase
    • AHUMAN NECESSITIES
    • A23FOODS OR FOODSTUFFS; TREATMENT THEREOF, NOT COVERED BY OTHER CLASSES
    • A23LFOODS, FOODSTUFFS OR NON-ALCOHOLIC BEVERAGES, NOT OTHERWISE PROVIDED FOR; PREPARATION OR TREATMENT THEREOF
    • A23L29/00Foods or foodstuffs containing additives; Preparation or treatment thereof
    • A23L29/30Foods or foodstuffs containing additives; Preparation or treatment thereof containing carbohydrate syrups; containing sugars; containing sugar alcohols, e.g. xylitol; containing starch hydrolysates, e.g. dextrin
    • A23L29/35Degradation products of starch, e.g. hydrolysates, dextrins; Enzymatically modified starches
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12NMICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14Hydrolases (3)
    • C12N9/24Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12NMICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14Hydrolases (3)
    • C12N9/24Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405Glucanases
    • C12N9/2408Glucanases acting on alpha -1,4-glucosidic bonds
    • C12N9/2411Amylases
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12NMICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14Hydrolases (3)
    • C12N9/24Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405Glucanases
    • C12N9/2408Glucanases acting on alpha -1,4-glucosidic bonds
    • C12N9/2411Amylases
    • C12N9/2428Glucan 1,4-alpha-glucosidase (3.2.1.3), i.e. glucoamylase
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12NMICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14Hydrolases (3)
    • C12N9/24Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405Glucanases
    • C12N9/2451Glucanases acting on alpha-1,6-glucosidic bonds
    • C12N9/2457Pullulanase (3.2.1.41)
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12PFERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P19/00Preparation of compounds containing saccharide radicals
    • C12P19/02Monosaccharides
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12YENZYMES
    • C12Y302/00Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01001Alpha-amylase (3.2.1.1)
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12YENZYMES
    • C12Y302/00Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01003Glucan 1,4-alpha-glucosidase (3.2.1.3), i.e. glucoamylase
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12YENZYMES
    • C12Y302/00Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01041Pullulanase (3.2.1.41)
    • CCHEMISTRY; METALLURGY
    • C13SUGAR INDUSTRY
    • C13KSACCHARIDES OBTAINED FROM NATURAL SOURCES OR BY HYDROLYSIS OF NATURALLY OCCURRING DISACCHARIDES, OLIGOSACCHARIDES OR POLYSACCHARIDES
    • C13K1/00Glucose; Glucose-containing syrups
    • C13K1/06Glucose; Glucose-containing syrups obtained by saccharification of starch or raw materials containing starch
    • YGENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y02TECHNOLOGIES OR APPLICATIONS FOR MITIGATION OR ADAPTATION AGAINST CLIMATE CHANGE
    • Y02EREDUCTION OF GREENHOUSE GAS [GHG] EMISSIONS, RELATED TO ENERGY GENERATION, TRANSMISSION OR DISTRIBUTION
    • Y02E50/00Technologies for the production of fuel of non-fossil origin
    • Y02E50/10Biofuels, e.g. bio-diesel

Landscapes

  • Chemical & Material Sciences (AREA)
  • Life Sciences & Earth Sciences (AREA)
  • Organic Chemistry (AREA)
  • Health & Medical Sciences (AREA)
  • Engineering & Computer Science (AREA)
  • Wood Science & Technology (AREA)
  • Zoology (AREA)
  • Bioinformatics & Cheminformatics (AREA)
  • Genetics & Genomics (AREA)
  • General Health & Medical Sciences (AREA)
  • Biochemistry (AREA)
  • General Engineering & Computer Science (AREA)
  • Microbiology (AREA)
  • Biotechnology (AREA)
  • Molecular Biology (AREA)
  • Biomedical Technology (AREA)
  • Medicinal Chemistry (AREA)
  • Chemical Kinetics & Catalysis (AREA)
  • General Chemical & Material Sciences (AREA)
  • Nutrition Science (AREA)
  • Food Science & Technology (AREA)
  • Polymers & Plastics (AREA)
  • Emergency Medicine (AREA)
  • Preparation Of Compounds By Using Micro-Organisms (AREA)
  • Enzymes And Modification Thereof (AREA)
  • Jellies, Jams, And Syrups (AREA)
  • Micro-Organisms Or Cultivation Processes Thereof (AREA)
JP2016550206A 2014-02-07 2015-02-06 グルコースシロップ生産用組成物 Active JP6591993B2 (ja)

Applications Claiming Priority (5)

Application Number Priority Date Filing Date Title
EP14154239.9 2014-02-07
EP14154239 2014-02-07
EP14195687.0 2014-12-01
EP14195687 2014-12-01
PCT/EP2015/052545 WO2015118123A1 (en) 2014-02-07 2015-02-06 Compositions for producing glucose syrups

Publications (3)

Publication Number Publication Date
JP2017505129A JP2017505129A (ja) 2017-02-16
JP2017505129A5 JP2017505129A5 (enExample) 2018-03-15
JP6591993B2 true JP6591993B2 (ja) 2019-10-16

Family

ID=52464388

Family Applications (1)

Application Number Title Priority Date Filing Date
JP2016550206A Active JP6591993B2 (ja) 2014-02-07 2015-02-06 グルコースシロップ生産用組成物

Country Status (10)

Country Link
US (3) US10337041B2 (enExample)
EP (2) EP3712240B1 (enExample)
JP (1) JP6591993B2 (enExample)
CN (1) CN105960176A (enExample)
BR (1) BR112016018075B1 (enExample)
CA (1) CA2937818C (enExample)
HU (1) HUE064397T2 (enExample)
MX (1) MX385596B (enExample)
RU (2) RU2019134734A (enExample)
WO (1) WO2015118123A1 (enExample)

Families Citing this family (4)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
FR3091703B1 (fr) 2019-01-11 2021-02-12 Fermentalg Procédé d’extraction de phycocyanines
FR3091640B1 (fr) 2019-01-11 2021-06-11 Fermentalg Procédé de purification de phycocyanines
FR3092586A1 (fr) 2019-02-08 2020-08-14 Fermentalg Procédé optimisé d’exploitation industrielle d’algues rouges unicellulaires
US20250311734A1 (en) 2022-05-14 2025-10-09 Novozymes A/S Compositions and Methods for Preventing, Treating, Suppressing and/or Eliminatting Phytopathogenic Infestations and Infections

Family Cites Families (46)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
JPS5534046A (en) 1978-09-01 1980-03-10 Cpc International Inc Novel glucoamyrase having excellent heat resistance and production
US4335208A (en) 1980-03-11 1982-06-15 Novo Industri A/S Saccharification of starch hydrolysates
NO840200L (no) 1983-01-28 1984-07-30 Cefus Corp Glukoamylase cdna.
US4587215A (en) 1984-06-25 1986-05-06 Uop Inc. Highly thermostable amyloglucosidase
JPS62126989A (ja) 1985-11-26 1987-06-09 Godo Shiyusei Kk コルテイシウム属担子菌の生産する酵素を用いた澱粉質の無蒸煮糖化法
JP2703598B2 (ja) 1987-09-04 1998-01-26 ノボ―ノルディスク アクティーゼルスカブ アスペルギルス菌中でのタンパク質生産物の生産方法およびアスペルギルス菌中で使用するためのプロモーター
US5162210A (en) 1990-06-29 1992-11-10 Iowa State University Research Foundation Process for enzymatic hydrolysis of starch to glucose
ATE183236T1 (de) 1992-12-28 1999-08-15 Genencor Int Pullulanase, mikroorganismen die sie produzieren, verfahren zur herstellung und ihre anwendung
JP4047379B2 (ja) 1995-02-03 2008-02-13 ノボザイムス アクティーゼルスカブ 予め決められた性質を有するα−アミラーゼ変異体のデザイン方法
EP0904360B1 (en) 1996-04-30 2013-07-31 Novozymes A/S alpha-AMYLASE MUTANTS
JP2000515377A (ja) * 1996-07-24 2000-11-21 アイオワ ステイト ユニヴァーシティ リサーチ ファウンデーション インコーポレイテッド pH最適値、基質特異性及び熱安定性を増大するためのグルコアミラーゼのタンパク質操作
AR014402A1 (es) 1997-11-26 2001-02-28 Novozymes As Glucoamilasa termoestable proceso para convertir almidon o almidon parcialmente hidrolizado en un jarabe que contiene dextrosa
KR100764528B1 (ko) 1998-07-15 2007-10-09 노보자임스 에이/에스 글루코아밀라제 변이체
AU5805700A (en) 1999-07-09 2001-01-30 Novozymes A/S Glucoamylase variant
EP1335982A2 (en) 2000-11-10 2003-08-20 Novozymes A/S Secondary liquefaction of starch in ethanol production
ES2278210T3 (es) 2002-12-17 2007-08-01 Novozymes A/S Alfa-amilasas termoestables.
JP2007526748A (ja) 2003-06-25 2007-09-20 ノボザイムス アクティーゼルスカブ 澱粉加工用酵素
ES2517245T3 (es) 2003-06-25 2014-11-03 Novozymes A/S Enzimas para el tratamiento de almidón
US7618795B2 (en) * 2003-06-25 2009-11-17 Novozymes A/S Starch process
CN1954072A (zh) * 2004-03-08 2007-04-25 辛根塔参与股份公司 自加工的植物和植物部分
CA2566252A1 (en) 2004-05-13 2005-12-01 Novozymes North America, Inc. A process of producing a fermentation product
ES2370768T3 (es) 2004-11-30 2011-12-22 Genencor International, Inc. Glucoamilasa de trichoderma reesei y homólogos de la misma.
ES2543131T3 (es) 2004-12-22 2015-08-14 Novozymes North America, Inc. Enzimas para tratamiento de almidón
WO2007144393A1 (en) * 2006-06-15 2007-12-21 Novozymes, Inc. Mashing process
MX2009003472A (es) 2006-10-10 2009-06-02 Danisco Us Inc Genencor Div Variantes de glucoamilasa con propiedades alteradas.
CN101049133B (zh) 2007-04-26 2011-07-06 华南理工大学 小麦b淀粉多酶法制取发酵用含氮糖浆的方法
PL2481796T3 (pl) 2007-10-09 2014-11-28 Danisco Us Inc Odmiany glukoamylazy
WO2009074650A2 (en) * 2007-12-12 2009-06-18 Novozymes A/S Brewing process
MX2010006016A (es) 2007-12-12 2010-07-01 Novozymes As Proceso de empastado de malta.
JP5827227B2 (ja) 2009-08-19 2015-12-02 デュポン ニュートリション バイオサイエンシーズ エーピーエスDuPont Nutrition Biosciences ApS グルコアミラーゼの変異体
WO2011022465A1 (en) 2009-08-19 2011-02-24 Danisco Us Inc. Combinatorial variants of glucoamylase with improved specific activity and/or thermostability
ES2534098T3 (es) 2009-11-30 2015-04-17 Novozymes A/S Polipéptidos que tienen actividad de glucoamilasa y polinucleótidos que codifican los mismos
CN104169417B (zh) * 2009-12-01 2020-05-08 诺维信公司 具有葡糖淀粉酶活性的多肽及其编码多核苷酸
WO2011127802A1 (en) 2010-04-14 2011-10-20 Novozymes A/S Polypeptides having glucoamylase activity and polynucleotides encoding same
WO2012064351A1 (en) 2010-11-08 2012-05-18 Novozymes A/S Polypeptides having glucoamylase activity and polynucleotides encoding same
KR101188113B1 (ko) 2010-12-23 2012-10-05 대상 주식회사 유동성이 향상된 쌀엿 제조방법
JP5928089B2 (ja) 2011-04-04 2016-06-01 国立大学法人弘前大学 イソマルターゼ
WO2013036526A1 (en) 2011-09-06 2013-03-14 Novozymes A/S Glucoamylase variants and polynucleotides encoding same
IN2014CN03467A (enExample) * 2011-10-11 2015-10-16 Novozymes North America Inc
FR2989892B1 (fr) 2012-04-26 2014-05-16 Oreal Composition cosmetique comprenant un oligosaccharide phosphoryle et un polysaccharide
MX2015002099A (es) * 2012-08-22 2015-05-11 Dupont Nutrition Biosci Aps Variantes que tienen actividad glucoamilasa.
CN121380029A (zh) * 2013-04-30 2026-01-23 诺维信公司 葡糖淀粉酶变体和编码它们的多核苷酸
CA2918685C (en) * 2013-08-30 2024-01-02 Novozymes A/S Enzyme composition and uses thereof
US10030237B2 (en) * 2014-01-22 2018-07-24 Novozymes A/S Pullulanase variants and polynucleotides encoding same
CN106795505A (zh) * 2014-10-23 2017-05-31 诺维信公司 葡糖淀粉酶变体和编码它们的多核苷酸
BR112017011275A2 (pt) * 2014-12-01 2018-04-03 Novozymes A/S composição, e, processo para a produção de xarope de glicose.

Also Published As

Publication number Publication date
EP3102051B1 (en) 2020-04-08
MX385596B (es) 2025-03-18
EP3712240B1 (en) 2023-09-27
JP2017505129A (ja) 2017-02-16
US20190256878A1 (en) 2019-08-22
US11136607B2 (en) 2021-10-05
US20170159090A1 (en) 2017-06-08
US11667938B2 (en) 2023-06-06
BR112016018075A2 (pt) 2018-02-20
MX2016010031A (es) 2016-11-15
CN105960176A (zh) 2016-09-21
CA2937818A1 (en) 2015-08-13
CA2937818C (en) 2023-08-01
HUE064397T2 (hu) 2024-03-28
RU2706297C2 (ru) 2019-11-15
US10337041B2 (en) 2019-07-02
BR112016018075B1 (pt) 2022-01-18
RU2019134734A (ru) 2019-11-18
EP3102051A1 (en) 2016-12-14
WO2015118123A1 (en) 2015-08-13
EP3712240A1 (en) 2020-09-23
RU2016135029A3 (enExample) 2018-10-16
RU2016135029A (ru) 2018-03-13
US20210395791A1 (en) 2021-12-23

Similar Documents

Publication Publication Date Title
US11667938B2 (en) Compositions for producing glucose syrups
US20090142818A1 (en) Process of producing a fermentation product
US20120252086A1 (en) Compositions Comprising Boosting Polypeptide And Starch Degrading Enzyme And Uses Thereof
US20110097779A1 (en) Processes for Producing Fermentation Products
US20080138871A1 (en) Fermentation Product Production Processes
US20200172946A1 (en) Production of Glucose Syrups
EP2326723B1 (en) Processes for producing fermentation products
US20160122442A1 (en) Process for Hydrolysis of Starch
CN105209627A (zh) 用于制备糖和糖浆的方法
WO2011100161A1 (en) Addition of alpha - glucosidase and cobalt for producing fermentation products from starch
US20080254518A1 (en) Liquefaction Processes
US20080009048A1 (en) Liquefaction of Starch Containing Material
US20080121227A1 (en) Liquefaction and Saccharification Process
US20070184150A1 (en) Liquefaction process

Legal Events

Date Code Title Description
A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20180205

A621 Written request for application examination

Free format text: JAPANESE INTERMEDIATE CODE: A621

Effective date: 20180205

A977 Report on retrieval

Free format text: JAPANESE INTERMEDIATE CODE: A971007

Effective date: 20181221

A131 Notification of reasons for refusal

Free format text: JAPANESE INTERMEDIATE CODE: A131

Effective date: 20190108

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20190408

A131 Notification of reasons for refusal

Free format text: JAPANESE INTERMEDIATE CODE: A131

Effective date: 20190507

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20190801

TRDD Decision of grant or rejection written
A01 Written decision to grant a patent or to grant a registration (utility model)

Free format text: JAPANESE INTERMEDIATE CODE: A01

Effective date: 20190820

A61 First payment of annual fees (during grant procedure)

Free format text: JAPANESE INTERMEDIATE CODE: A61

Effective date: 20190919

R150 Certificate of patent or registration of utility model

Ref document number: 6591993

Country of ref document: JP

Free format text: JAPANESE INTERMEDIATE CODE: R150

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250