JP6591993B2 - グルコースシロップ生産用組成物 - Google Patents

グルコースシロップ生産用組成物 Download PDF

Info

Publication number
JP6591993B2
JP6591993B2 JP2016550206A JP2016550206A JP6591993B2 JP 6591993 B2 JP6591993 B2 JP 6591993B2 JP 2016550206 A JP2016550206 A JP 2016550206A JP 2016550206 A JP2016550206 A JP 2016550206A JP 6591993 B2 JP6591993 B2 JP 6591993B2
Authority
JP
Japan
Prior art keywords
seq
amylase
pullulanase
glucoamylase
mature polypeptide
Prior art date
Legal status (The legal status is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the status listed.)
Active
Application number
JP2016550206A
Other languages
English (en)
Japanese (ja)
Other versions
JP2017505129A (ja
JP2017505129A5 (enExample
Inventor
志朗 福山
志朗 福山
圭一 綾部
圭一 綾部
Original Assignee
ノボザイムス アクティーゼルスカブ
ノボザイムス アクティーゼルスカブ
Priority date (The priority date is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the date listed.)
Filing date
Publication date
Application filed by ノボザイムス アクティーゼルスカブ, ノボザイムス アクティーゼルスカブ filed Critical ノボザイムス アクティーゼルスカブ
Publication of JP2017505129A publication Critical patent/JP2017505129A/ja
Publication of JP2017505129A5 publication Critical patent/JP2017505129A5/ja
Application granted granted Critical
Publication of JP6591993B2 publication Critical patent/JP6591993B2/ja
Active legal-status Critical Current
Anticipated expiration legal-status Critical

Links

Classifications

    • A—HUMAN NECESSITIES
    • A23—FOODS OR FOODSTUFFS; TREATMENT THEREOF, NOT COVERED BY OTHER CLASSES
    • A23L—FOODS, FOODSTUFFS OR NON-ALCOHOLIC BEVERAGES, NOT OTHERWISE PROVIDED FOR; PREPARATION OR TREATMENT THEREOF
    • A23L29/00—Foods or foodstuffs containing additives; Preparation or treatment thereof
    • A23L29/30—Foods or foodstuffs containing additives; Preparation or treatment thereof containing carbohydrate syrups; containing sugars; containing sugar alcohols, e.g. xylitol; containing starch hydrolysates, e.g. dextrin
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12P—FERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P19/00—Preparation of compounds containing saccharide radicals
    • C12P19/14—Preparation of compounds containing saccharide radicals produced by the action of a carbohydrase (EC 3.2.x), e.g. by alpha-amylase, e.g. by cellulase, hemicellulase
    • A—HUMAN NECESSITIES
    • A23—FOODS OR FOODSTUFFS; TREATMENT THEREOF, NOT COVERED BY OTHER CLASSES
    • A23L—FOODS, FOODSTUFFS OR NON-ALCOHOLIC BEVERAGES, NOT OTHERWISE PROVIDED FOR; PREPARATION OR TREATMENT THEREOF
    • A23L29/00—Foods or foodstuffs containing additives; Preparation or treatment thereof
    • A23L29/30—Foods or foodstuffs containing additives; Preparation or treatment thereof containing carbohydrate syrups; containing sugars; containing sugar alcohols, e.g. xylitol; containing starch hydrolysates, e.g. dextrin
    • A23L29/35—Degradation products of starch, e.g. hydrolysates, dextrins; Enzymatically modified starches
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405—Glucanases
    • C12N9/2408—Glucanases acting on alpha -1,4-glucosidic bonds
    • C12N9/2411—Amylases
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405—Glucanases
    • C12N9/2408—Glucanases acting on alpha -1,4-glucosidic bonds
    • C12N9/2411—Amylases
    • C12N9/2428—Glucan 1,4-alpha-glucosidase (3.2.1.3), i.e. glucoamylase
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405—Glucanases
    • C12N9/2451—Glucanases acting on alpha-1,6-glucosidic bonds
    • C12N9/2457—Pullulanase (3.2.1.41)
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12P—FERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P19/00—Preparation of compounds containing saccharide radicals
    • C12P19/02—Monosaccharides
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12Y—ENZYMES
    • C12Y302/00—Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01—Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01001—Alpha-amylase (3.2.1.1)
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12Y—ENZYMES
    • C12Y302/00—Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01—Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01003—Glucan 1,4-alpha-glucosidase (3.2.1.3), i.e. glucoamylase
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12Y—ENZYMES
    • C12Y302/00—Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01—Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01041—Pullulanase (3.2.1.41)
    • C—CHEMISTRY; METALLURGY
    • C13—SUGAR INDUSTRY
    • C13K—SACCHARIDES OBTAINED FROM NATURAL SOURCES OR BY HYDROLYSIS OF NATURALLY OCCURRING DISACCHARIDES, OLIGOSACCHARIDES OR POLYSACCHARIDES
    • C13K1/00—Glucose; Glucose-containing syrups
    • C13K1/06—Glucose; Glucose-containing syrups obtained by saccharification of starch or raw materials containing starch
    • Y—GENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y02—TECHNOLOGIES OR APPLICATIONS FOR MITIGATION OR ADAPTATION AGAINST CLIMATE CHANGE
    • Y02E—REDUCTION OF GREENHOUSE GAS [GHG] EMISSIONS, RELATED TO ENERGY GENERATION, TRANSMISSION OR DISTRIBUTION
    • Y02E50/00—Technologies for the production of fuel of non-fossil origin
    • Y02E50/10—Biofuels, e.g. bio-diesel

Landscapes

  • Chemical & Material Sciences (AREA)
  • Life Sciences & Earth Sciences (AREA)
  • Organic Chemistry (AREA)
  • Health & Medical Sciences (AREA)
  • Engineering & Computer Science (AREA)
  • Wood Science & Technology (AREA)
  • Zoology (AREA)
  • Bioinformatics & Cheminformatics (AREA)
  • Genetics & Genomics (AREA)
  • General Health & Medical Sciences (AREA)
  • Biochemistry (AREA)
  • General Engineering & Computer Science (AREA)
  • Microbiology (AREA)
  • Biotechnology (AREA)
  • Molecular Biology (AREA)
  • Medicinal Chemistry (AREA)
  • Biomedical Technology (AREA)
  • General Chemical & Material Sciences (AREA)
  • Chemical Kinetics & Catalysis (AREA)
  • Nutrition Science (AREA)
  • Food Science & Technology (AREA)
  • Polymers & Plastics (AREA)
  • Emergency Medicine (AREA)
  • Preparation Of Compounds By Using Micro-Organisms (AREA)
  • Enzymes And Modification Thereof (AREA)
  • Jellies, Jams, And Syrups (AREA)
  • Micro-Organisms Or Cultivation Processes Thereof (AREA)
JP2016550206A 2014-02-07 2015-02-06 グルコースシロップ生産用組成物 Active JP6591993B2 (ja)

Applications Claiming Priority (5)

Application Number Priority Date Filing Date Title
EP14154239 2014-02-07
EP14154239.9 2014-02-07
EP14195687 2014-12-01
EP14195687.0 2014-12-01
PCT/EP2015/052545 WO2015118123A1 (en) 2014-02-07 2015-02-06 Compositions for producing glucose syrups

Publications (3)

Publication Number Publication Date
JP2017505129A JP2017505129A (ja) 2017-02-16
JP2017505129A5 JP2017505129A5 (enExample) 2018-03-15
JP6591993B2 true JP6591993B2 (ja) 2019-10-16

Family

ID=52464388

Family Applications (1)

Application Number Title Priority Date Filing Date
JP2016550206A Active JP6591993B2 (ja) 2014-02-07 2015-02-06 グルコースシロップ生産用組成物

Country Status (10)

Country Link
US (3) US10337041B2 (enExample)
EP (2) EP3102051B1 (enExample)
JP (1) JP6591993B2 (enExample)
CN (1) CN105960176A (enExample)
BR (1) BR112016018075B1 (enExample)
CA (1) CA2937818C (enExample)
HU (1) HUE064397T2 (enExample)
MX (1) MX385596B (enExample)
RU (2) RU2019134734A (enExample)
WO (1) WO2015118123A1 (enExample)

Families Citing this family (5)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
FR3091703B1 (fr) 2019-01-11 2021-02-12 Fermentalg Procédé d’extraction de phycocyanines
FR3091640B1 (fr) 2019-01-11 2021-06-11 Fermentalg Procédé de purification de phycocyanines
FR3092586A1 (fr) 2019-02-08 2020-08-14 Fermentalg Procédé optimisé d’exploitation industrielle d’algues rouges unicellulaires
WO2023225459A2 (en) 2022-05-14 2023-11-23 Novozymes A/S Compositions and methods for preventing, treating, supressing and/or eliminating phytopathogenic infestations and infections
WO2026145739A1 (zh) * 2024-12-31 2026-07-09 南京百斯杰生物工程有限公司 α-淀粉酶突变体及其制备方法与应用

Family Cites Families (46)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
JPS5534046A (en) 1978-09-01 1980-03-10 Cpc International Inc Novel glucoamyrase having excellent heat resistance and production
US4335208A (en) 1980-03-11 1982-06-15 Novo Industri A/S Saccharification of starch hydrolysates
NO840200L (no) 1983-01-28 1984-07-30 Cefus Corp Glukoamylase cdna.
US4587215A (en) 1984-06-25 1986-05-06 Uop Inc. Highly thermostable amyloglucosidase
JPS62126989A (ja) 1985-11-26 1987-06-09 Godo Shiyusei Kk コルテイシウム属担子菌の生産する酵素を用いた澱粉質の無蒸煮糖化法
DE3886221T3 (de) 1987-09-04 2000-12-21 Novo Nordisk A/S, Bagsvaerd VERFAHREN ZUR HERSTELLUNG VON PROTEINPRODUKTEN IN -i(ASPERGILLUS) UND PROMOTOREN ZUR VERWENDUNG IN -i(ASPERGILLUS).
US5162210A (en) 1990-06-29 1992-11-10 Iowa State University Research Foundation Process for enzymatic hydrolysis of starch to glucose
ATE183236T1 (de) 1992-12-28 1999-08-15 Genencor Int Pullulanase, mikroorganismen die sie produzieren, verfahren zur herstellung und ihre anwendung
MX9705906A (es) 1995-02-03 1997-10-31 Novo Nordisk As Un metodo para diseñar mutantes de alfa-amilasa con propiedades predeterminadas.
AU2692897A (en) 1996-04-30 1997-11-19 Novo Nordisk A/S Alpha-amylase mutants
CN1238009A (zh) * 1996-07-24 1999-12-08 衣阿华州立大学研究基金会股份有限公司 提高葡糖淀粉酶pH最适点、底物专一性和热稳定性的蛋白质工程方法
DK1032654T3 (da) 1997-11-26 2009-05-11 Novozymes As Thermostabil glucoamylase
CA2331340A1 (en) 1998-07-15 2000-01-27 Novozymes A/S Glucoamylase variants
AR029759A1 (es) 1999-07-09 2003-07-16 Novozymes As Variante de glucoamilasa original, una construccion de adn, un vector de expresion recombinante, una celula, un proceso para convertir almidon, uso de una variante de glucoamilasa para un proceso de fermentacion y metodo para mejorar la estabilidad termica.
WO2002038787A2 (en) * 2000-11-10 2002-05-16 Novozymes A/S Secondary liquefaction of starch in ethanol production
DE60310264T2 (de) 2002-12-17 2007-07-05 Novozymes A/S Thermostabile alpha-amylase
US7618795B2 (en) * 2003-06-25 2009-11-17 Novozymes A/S Starch process
MXPA06000212A (es) 2003-06-25 2006-03-21 Novozymes As Enzimas para procesar almidon.
ATE510925T1 (de) * 2003-06-25 2011-06-15 Novozymes As Stärkehydrolyseverfahren
BRPI0418622B1 (pt) * 2004-03-08 2020-01-28 Syngenta Participations Ag polinucleotídeo isolado, cassete de expressão e método para preparar açúcar fermentável, monossacarídeo ou oligossacarídeo
MXPA06013130A (es) * 2004-05-13 2007-04-19 Novozymes North America Inc Proceso para producir un producto de fermentacion.
WO2006060062A2 (en) * 2004-11-30 2006-06-08 Genencor International, Inc. Trichoderma reesei glucoamylase and homologs thereof
MX2007007107A (es) * 2004-12-22 2007-08-08 Novozymes North America Inc Polipeptidos que tienen actividad de glucoamilasa y polinucleotidos que los codifican.
WO2007144393A1 (en) 2006-06-15 2007-12-21 Novozymes, Inc. Mashing process
EP2061882A2 (en) 2006-10-10 2009-05-27 Danisco US, INC., Genencor Division Glucoamylase variants with altered properties
CN101049133B (zh) 2007-04-26 2011-07-06 华南理工大学 小麦b淀粉多酶法制取发酵用含氮糖浆的方法
PL2481796T3 (pl) 2007-10-09 2014-11-28 Danisco Us Inc Odmiany glukoamylazy
DK2222830T3 (en) 2007-12-12 2015-11-16 Novozymes Inc A method of mashing
WO2009074650A2 (en) * 2007-12-12 2009-06-18 Novozymes A/S Brewing process
AR077942A1 (es) 2009-08-19 2011-10-05 Danisco Variantes de glucoamilasa
EP2467475A1 (en) 2009-08-19 2012-06-27 Danisco US Inc. Combinatorial variants of glucoamylase with improved specific activity and/or thermostability
CA2782154C (en) 2009-11-30 2018-10-16 Novozymes A/S Polypeptides having glucoamylase activity and polynucleotides encoding same
CA2782036C (en) * 2009-12-01 2019-01-15 Novozymes A/S Polypeptides having glucoamylase activity and polynucleotides encoding same
CA2795806C (en) * 2010-04-14 2019-01-08 Novozymes A/S Polypeptides having glucoamylase activity and polynucleotides encoding same
DK2638154T3 (da) 2010-11-08 2017-01-02 Novozymes As Polypeptider med glucoamylaseaktivitet og polynukleotider, der koder for dem
KR101188113B1 (ko) * 2010-12-23 2012-10-05 대상 주식회사 유동성이 향상된 쌀엿 제조방법
JP5928089B2 (ja) * 2011-04-04 2016-06-01 国立大学法人弘前大学 イソマルターゼ
CN103930543B (zh) 2011-09-06 2020-11-27 诺维信公司 葡糖淀粉酶变体和编码它们的多核苷酸
EP2766490B1 (en) * 2011-10-11 2017-04-12 Novozymes North America, Inc. Liquefaction process for producing fermentation products
FR2989892B1 (fr) * 2012-04-26 2014-05-16 Oreal Composition cosmetique comprenant un oligosaccharide phosphoryle et un polysaccharide
MX2015002099A (es) * 2012-08-22 2015-05-11 Dupont Nutrition Biosci Aps Variantes que tienen actividad glucoamilasa.
EP3372680B1 (en) * 2013-04-30 2020-11-11 Novozymes A/S Glucoamylase variants and polynucleotides encoding same
EP4717767A2 (en) * 2013-08-30 2026-04-01 Novozymes A/S Enzyme composition and uses thereof
EP3097192B1 (en) * 2014-01-22 2018-07-25 Novozymes A/S Pullulanase variants and polynucleotides encoding same
WO2016062875A2 (en) * 2014-10-23 2016-04-28 Novozymes A/S Glucoamylase variants and polynucleotides encoding same
BR112017011275A2 (pt) * 2014-12-01 2018-04-03 Novozymes A/S composição, e, processo para a produção de xarope de glicose.

Also Published As

Publication number Publication date
MX2016010031A (es) 2016-11-15
US20210395791A1 (en) 2021-12-23
RU2019134734A (ru) 2019-11-18
BR112016018075B1 (pt) 2022-01-18
CN105960176A (zh) 2016-09-21
WO2015118123A1 (en) 2015-08-13
MX385596B (es) 2025-03-18
EP3102051B1 (en) 2020-04-08
CA2937818C (en) 2023-08-01
US20170159090A1 (en) 2017-06-08
JP2017505129A (ja) 2017-02-16
CA2937818A1 (en) 2015-08-13
US10337041B2 (en) 2019-07-02
HUE064397T2 (hu) 2024-03-28
US11667938B2 (en) 2023-06-06
US11136607B2 (en) 2021-10-05
US20190256878A1 (en) 2019-08-22
EP3102051A1 (en) 2016-12-14
EP3712240A1 (en) 2020-09-23
RU2706297C2 (ru) 2019-11-15
BR112016018075A2 (pt) 2018-02-20
EP3712240B1 (en) 2023-09-27
RU2016135029A3 (enExample) 2018-10-16
RU2016135029A (ru) 2018-03-13

Similar Documents

Publication Publication Date Title
US11667938B2 (en) Compositions for producing glucose syrups
US20090142818A1 (en) Process of producing a fermentation product
US20120252086A1 (en) Compositions Comprising Boosting Polypeptide And Starch Degrading Enzyme And Uses Thereof
US20110097779A1 (en) Processes for Producing Fermentation Products
US20080138871A1 (en) Fermentation Product Production Processes
US20200172946A1 (en) Production of Glucose Syrups
EP2326723B1 (en) Processes for producing fermentation products
US20160122442A1 (en) Process for Hydrolysis of Starch
US20080121227A1 (en) Liquefaction and Saccharification Process
CN105209627A (zh) 用于制备糖和糖浆的方法
WO2011100161A1 (en) Addition of alpha - glucosidase and cobalt for producing fermentation products from starch
US20080254518A1 (en) Liquefaction Processes
US20080009048A1 (en) Liquefaction of Starch Containing Material
US20070184150A1 (en) Liquefaction process

Legal Events

Date Code Title Description
A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20180205

A621 Written request for application examination

Free format text: JAPANESE INTERMEDIATE CODE: A621

Effective date: 20180205

A977 Report on retrieval

Free format text: JAPANESE INTERMEDIATE CODE: A971007

Effective date: 20181221

A131 Notification of reasons for refusal

Free format text: JAPANESE INTERMEDIATE CODE: A131

Effective date: 20190108

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20190408

A131 Notification of reasons for refusal

Free format text: JAPANESE INTERMEDIATE CODE: A131

Effective date: 20190507

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20190801

TRDD Decision of grant or rejection written
A01 Written decision to grant a patent or to grant a registration (utility model)

Free format text: JAPANESE INTERMEDIATE CODE: A01

Effective date: 20190820

A61 First payment of annual fees (during grant procedure)

Free format text: JAPANESE INTERMEDIATE CODE: A61

Effective date: 20190919

R150 Certificate of patent or registration of utility model

Ref document number: 6591993

Country of ref document: JP

Free format text: JAPANESE INTERMEDIATE CODE: R150

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250