CN101378651B - 氮源高效利用的单子叶植物 - Google Patents
氮源高效利用的单子叶植物 Download PDFInfo
- Publication number
- CN101378651B CN101378651B CN2006800487181A CN200680048718A CN101378651B CN 101378651 B CN101378651 B CN 101378651B CN 2006800487181 A CN2006800487181 A CN 2006800487181A CN 200680048718 A CN200680048718 A CN 200680048718A CN 101378651 B CN101378651 B CN 101378651B
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- China
- Prior art keywords
- plant
- promotor
- transgenic
- paddy rice
- nucleotide
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- Legal status (The legal status is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the status listed.)
- Expired - Fee Related
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- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8241—Phenotypically and genetically modified plants via recombinant DNA technology
- C12N15/8261—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield
- C12N15/8262—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield involving plant development
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- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
- C12N9/10—Transferases (2.)
- C12N9/1096—Transferases (2.) transferring nitrogenous groups (2.6)
-
- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8216—Methods for controlling, regulating or enhancing expression of transgenes in plant cells
- C12N15/8222—Developmentally regulated expression systems, tissue, organ specific, temporal or spatial regulation
- C12N15/8223—Vegetative tissue-specific promoters
- C12N15/8227—Root-specific
-
- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8241—Phenotypically and genetically modified plants via recombinant DNA technology
- C12N15/8242—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits
- C12N15/8243—Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits involving biosynthetic or metabolic pathways, i.e. metabolic engineering, e.g. nicotine, caffeine
-
- C—CHEMISTRY; METALLURGY
- C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
- C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
- C12N15/00—Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
- C12N15/09—Recombinant DNA-technology
- C12N15/63—Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
- C12N15/79—Vectors or expression systems specially adapted for eukaryotic hosts
- C12N15/82—Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
- C12N15/8241—Phenotypically and genetically modified plants via recombinant DNA technology
- C12N15/8261—Phenotypically and genetically modified plants via recombinant DNA technology with agronomic (input) traits, e.g. crop yield
-
- Y—GENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
- Y02—TECHNOLOGIES OR APPLICATIONS FOR MITIGATION OR ADAPTATION AGAINST CLIMATE CHANGE
- Y02A—TECHNOLOGIES FOR ADAPTATION TO CLIMATE CHANGE
- Y02A40/00—Adaptation technologies in agriculture, forestry, livestock or agroalimentary production
- Y02A40/10—Adaptation technologies in agriculture, forestry, livestock or agroalimentary production in agriculture
- Y02A40/146—Genetically Modified [GMO] plants, e.g. transgenic plants
Landscapes
- Health & Medical Sciences (AREA)
- Genetics & Genomics (AREA)
- Life Sciences & Earth Sciences (AREA)
- Engineering & Computer Science (AREA)
- Chemical & Material Sciences (AREA)
- Zoology (AREA)
- Organic Chemistry (AREA)
- Biomedical Technology (AREA)
- Biotechnology (AREA)
- Bioinformatics & Cheminformatics (AREA)
- Wood Science & Technology (AREA)
- General Engineering & Computer Science (AREA)
- Molecular Biology (AREA)
- Microbiology (AREA)
- Biochemistry (AREA)
- General Health & Medical Sciences (AREA)
- Cell Biology (AREA)
- Physics & Mathematics (AREA)
- Biophysics (AREA)
- Plant Pathology (AREA)
- Medicinal Chemistry (AREA)
- Nutrition Science (AREA)
- Breeding Of Plants And Reproduction By Means Of Culturing (AREA)
- Micro-Organisms Or Cultivation Processes Thereof (AREA)
- Preparation Of Compounds By Using Micro-Organisms (AREA)
- Enzymes And Modification Thereof (AREA)
Applications Claiming Priority (3)
Application Number | Priority Date | Filing Date | Title |
---|---|---|---|
US75381805P | 2005-12-23 | 2005-12-23 | |
US60/753,818 | 2005-12-23 | ||
PCT/US2006/049241 WO2007076115A2 (en) | 2005-12-23 | 2006-12-21 | Nitrogen-efficient monocot plants |
Publications (2)
Publication Number | Publication Date |
---|---|
CN101378651A CN101378651A (zh) | 2009-03-04 |
CN101378651B true CN101378651B (zh) | 2012-03-14 |
Family
ID=38218711
Family Applications (1)
Application Number | Title | Priority Date | Filing Date |
---|---|---|---|
CN2006800487181A Expired - Fee Related CN101378651B (zh) | 2005-12-23 | 2006-12-21 | 氮源高效利用的单子叶植物 |
Country Status (9)
Country | Link |
---|---|
US (2) | US8288611B2 (pt) |
EP (1) | EP1983819A4 (pt) |
CN (1) | CN101378651B (pt) |
AR (2) | AR059119A1 (pt) |
AU (1) | AU2006330817B2 (pt) |
BR (1) | BRPI0620312A2 (pt) |
CA (1) | CA2634925C (pt) |
WO (1) | WO2007076115A2 (pt) |
ZA (1) | ZA200805007B (pt) |
Families Citing this family (41)
Publication number | Priority date | Publication date | Assignee | Title |
---|---|---|---|---|
US6084153A (en) * | 1996-02-14 | 2000-07-04 | The Governors Of The University Of Alberta | Plants having enhanced nitrogen assimilation/metabolism |
WO2007075925A2 (en) * | 2005-12-23 | 2007-07-05 | Arcadia Biosciences, Inc. | Promoter sequence obtained from rice and methods of use |
AR059119A1 (es) | 2005-12-23 | 2008-03-12 | Arcadia Biosciences Inc | Utilizacion eficaz de nitrogeno en plantas monocotiledoneas |
CA2649148C (en) | 2006-04-13 | 2016-08-23 | Chugai Seiyaku Kabushiki Kaisha | Taurine transporter gene |
JP5635260B2 (ja) | 2007-03-15 | 2014-12-03 | 中外製薬株式会社 | ポリペプチドの製造方法 |
AR070640A1 (es) * | 2007-07-20 | 2010-04-28 | Cropdesign Nv | Plantas que tienen rasgos mejorados relacionados con el rendimiento y un metodo para obtenerlas |
ES2440265T3 (es) * | 2007-07-20 | 2014-01-28 | Basf Plant Science Gmbh | Plantas que tienen un aumento de características relacionadas con el rendimiento y un método para elaboración de las mismas |
RU2487168C2 (ru) | 2007-08-07 | 2013-07-10 | Чугаи Сейяку Кабусики Кайся | Способ получения гетерологичных белков |
WO2009039517A1 (en) * | 2007-09-21 | 2009-03-26 | Arcadia Biosciences, Inc. | Energy efficient and greenhouse gas efficient biofuel |
JP5337043B2 (ja) * | 2007-10-15 | 2013-11-06 | 中外製薬株式会社 | 異種タンパク質を高生産する細胞の作製方法 |
EP2213746B1 (en) * | 2007-10-24 | 2015-07-15 | Chugai Seiyaku Kabushiki Kaisha | Cell for use in the production of exogenous protein, and production process using the cell |
WO2009146015A2 (en) * | 2008-03-31 | 2009-12-03 | Ceres, Inc. | Promoter, promoter control elements, and combinations, and uses thereof |
WO2009149304A2 (en) * | 2008-06-04 | 2009-12-10 | Edenspace Systems Corporation | Plant gene regulatory elements |
US20110004961A1 (en) * | 2008-08-29 | 2011-01-06 | Unkefer Pat J | Plant glutamine phenylpyruvate transaminase gene and transgenic plants carrying same |
US20110030089A1 (en) | 2008-08-29 | 2011-02-03 | Los Alamos National Security, Llc | Transgenic plants with enhanced growth characteristics |
US20110030104A1 (en) | 2008-08-29 | 2011-02-03 | Los Alamos National Security, Llc | Nucleic acids encoding plant glutamine phenylpyruvate transaminase (GPT) and uses thereof |
EP2511374A1 (en) * | 2009-03-27 | 2012-10-17 | E. I. du Pont de Nemours and Company | Plants having altered agronomic characteristics under nitrogen limiting conditions and related constructs and methods involving genes encoding SNF2 domain-containing polypeptides |
UA108071C2 (uk) * | 2009-04-14 | 2015-03-25 | Піонер Хай-Бред Інтернешнл, Інк. | Спосіб поліпшення витривалості до нестачі азоту у рослини |
JP5715050B2 (ja) | 2009-04-22 | 2015-05-07 | 中外製薬株式会社 | 異種タンパク質を高生産する細胞の作製方法 |
US20110061132A1 (en) | 2009-08-20 | 2011-03-10 | Pioneer Hi-Bred International, Inc. | Functional expression of yeast nitrate transporter (ynt1) in maize to improve nitrate uptake |
MA33933B1 (fr) | 2010-01-22 | 2013-01-02 | Bayer Ip Gmbh | Combinaisons de principes actifs acaricides et/ou insecticides |
CN101942512B (zh) * | 2010-08-25 | 2012-10-10 | 中国农业科学院作物科学研究所 | 玉米低氮逆境下高籽粒数目优异等位基因功能分子标记开发与应用 |
WO2012049663A1 (fr) | 2010-10-15 | 2012-04-19 | Genoplante-Valor | Obtention de plantes ayant une tolérance améliorée à un déficit hydrique |
FR2966162B1 (fr) | 2010-10-15 | 2014-08-29 | Genoplante Valor | Obtention de plantes ayant une tolerance amelioree a un deficit hydrique |
UA109460C2 (uk) | 2010-11-02 | 2015-08-25 | Байєр Інтелекчуал Проперті Гмбх | N-гетарилметилпіразолілкарбоксаміди |
JP5860471B2 (ja) | 2010-11-15 | 2016-02-16 | バイエル・インテレクチュアル・プロパティ・ゲゼルシャフト・ミット・ベシュレンクテル・ハフツングBayer Intellectual Property GmbH | N−アリールピラゾール(チオ)カルボキサミド類 |
EP2658853A1 (en) | 2010-12-29 | 2013-11-06 | Bayer Intellectual Property GmbH | Fungicide hydroximoyl-tetrazole derivatives |
FR2976950A1 (fr) | 2011-06-24 | 2012-12-28 | Genoplante Valor | Obtention de plantes ayant une tolerance amelioree a un stress hydrique |
CN103717076B (zh) | 2011-08-10 | 2016-04-13 | 拜耳知识产权股份有限公司 | 含有特定特特拉姆酸衍生物的活性化合物组合物 |
UA116086C2 (uk) * | 2011-09-07 | 2018-02-12 | Байєр Кропсаєнс Аг | Трансгенна однодольна рослина з підвищеною ефективністю використання азоту |
FR2994793A1 (fr) | 2012-08-30 | 2014-03-07 | Agronomique Inst Nat Rech | Utilisation d'un recepteur kinase a motifs lysm pour ameliorer la reponse des plantes aux lipochitooligosaccharides. |
US9695436B2 (en) * | 2013-03-06 | 2017-07-04 | Bayer Cropscience Lp | Plants having enhanced nitrogen use efficiency and methods of producing same |
CN103392591B (zh) * | 2013-07-05 | 2016-08-10 | 安徽省农业科学院水稻研究所 | 一种水稻高产且氮素高效利用的新品种选育方法 |
CN109112138B (zh) * | 2017-12-01 | 2020-12-08 | 华中农业大学 | 一种调控水稻理想株型的基因OsVAS1 |
AR118786A1 (es) | 2018-08-13 | 2021-11-03 | Univ Aarhus | Plantas alteradas genéticamente que expresan receptores heterólogos que reconocen lipoquitooligosacáridos |
AR118785A1 (es) | 2018-08-13 | 2021-11-03 | Univ Aarhus | RECEPTORES DE LysM GENÉTICAMENTE ALTERADOS CON ESPECIFICIDAD Y AFINIDAD AGONISTAS ALTERADAS |
CA3121296A1 (en) | 2018-12-06 | 2020-06-11 | Wageningen Universiteit | Methods of genetically altering a plant nin-gene to be responsive to cytokinin |
CA3133153A1 (en) | 2019-03-21 | 2020-09-24 | University Of Essex Enterprises Limited | Methods of enhancing biomass in a plant through stimulation of rubp regeneration and electron transport |
AU2021274836A1 (en) | 2020-05-19 | 2023-01-05 | Aarhus Universitet | Lysm receptor motifs |
WO2022251428A2 (en) | 2021-05-26 | 2022-12-01 | The Board Of Trustees Of The University Of Illinois | C4 plants with increased photosynthetic efficiency |
US20230323480A1 (en) | 2022-04-11 | 2023-10-12 | The Regents Of The University Of California | Methods of screening for plant gain of function mutations and compositions therefor |
Citations (2)
Publication number | Priority date | Publication date | Assignee | Title |
---|---|---|---|---|
WO1997030163A1 (en) * | 1996-02-14 | 1997-08-21 | The Governors Of The University Of Alberta | Plants having enhanced nitrogen assimilation/metabolism |
WO2001055433A2 (en) * | 2000-01-28 | 2001-08-02 | The Governors Of The University Of Alberta | Tissue-specific expression of target genes in plants |
Family Cites Families (17)
Publication number | Priority date | Publication date | Assignee | Title |
---|---|---|---|---|
DE3719053A1 (de) | 1987-06-06 | 1988-12-15 | Hoechst Ag | Verbesserte nutzung von pflanzenverwertbarem stickstoff durch kulturpflanzen mit ueberexpression der glutaminsynthetase |
US5955651A (en) * | 1989-05-03 | 1999-09-21 | New York University | Transgenic plants that exhibit enhanced nitrogen assimilation |
US5256558A (en) * | 1989-05-03 | 1993-10-26 | The Trustees Of Rockefeller University | Gene encoding plant asparagine synthetase |
EP0491780A4 (en) | 1989-09-15 | 1992-11-19 | The General Hospital Corporation | A membrane associated protein from plants which has nitrate reductase activity |
US5399680A (en) | 1991-05-22 | 1995-03-21 | The Salk Institute For Biological Studies | Rice chitinase promoter |
JPH05199877A (ja) | 1991-10-03 | 1993-08-10 | Sumitomo Chem Co Ltd | バレイショおよびイネの誘導型植物防御遺伝子の制御領域、その用途およびアッセイ法 |
ES2093578T1 (es) | 1993-10-06 | 1997-01-01 | Univ New York | Plantas transgenicas que presentan asimilacion mejorada del nitrogeno. |
US5750399A (en) * | 1994-11-14 | 1998-05-12 | The Samuel Roberts Noble Foundation, Inc. | Isoflavone reductase promoter |
US7390937B2 (en) * | 1996-02-14 | 2008-06-24 | The Governors Of The University Of Alberta | Plants with enhanced levels of nitrogen utilization proteins in their root epidermis and uses thereof |
US6084153A (en) * | 1996-02-14 | 2000-07-04 | The Governors Of The University Of Alberta | Plants having enhanced nitrogen assimilation/metabolism |
US6080913A (en) * | 1996-09-25 | 2000-06-27 | Pioneer Hi-Bred International, Inc. | Binary methods of increasing accumulation of essential amino acids in seeds |
US20040116682A1 (en) * | 1998-03-06 | 2004-06-17 | Nordine Cheikh | Nucleic acid molecules and other molecules associated with the carbon assimilation pathway |
US7365185B2 (en) * | 2000-07-19 | 2008-04-29 | Monsanto Technology Llc | Genomic plant sequences and uses thereof |
EP1402037A1 (en) | 2001-06-22 | 2004-03-31 | Syngenta Participations AG | Plant genes involved in defense against pathogens |
WO2004003146A2 (en) * | 2002-06-28 | 2004-01-08 | Paradigm Genetics, Inc. | Method for improving plant traits by altering the activity of plant g-protein alpha and beta subunits |
AR059119A1 (es) | 2005-12-23 | 2008-03-12 | Arcadia Biosciences Inc | Utilizacion eficaz de nitrogeno en plantas monocotiledoneas |
WO2007075925A2 (en) * | 2005-12-23 | 2007-07-05 | Arcadia Biosciences, Inc. | Promoter sequence obtained from rice and methods of use |
-
2006
- 2006-12-21 AR ARP060105745A patent/AR059119A1/es not_active Application Discontinuation
- 2006-12-21 EP EP06848873A patent/EP1983819A4/en not_active Ceased
- 2006-12-21 US US11/644,321 patent/US8288611B2/en active Active
- 2006-12-21 BR BRPI0620312-4A patent/BRPI0620312A2/pt not_active Application Discontinuation
- 2006-12-21 CN CN2006800487181A patent/CN101378651B/zh not_active Expired - Fee Related
- 2006-12-21 WO PCT/US2006/049241 patent/WO2007076115A2/en active Application Filing
- 2006-12-21 AU AU2006330817A patent/AU2006330817B2/en not_active Ceased
- 2006-12-21 CA CA2634925A patent/CA2634925C/en not_active Expired - Fee Related
-
2008
- 2008-06-09 ZA ZA200805007A patent/ZA200805007B/xx unknown
-
2012
- 2012-09-18 US US13/622,311 patent/US8642840B2/en not_active Expired - Fee Related
-
2013
- 2013-11-04 AR ARP130104024A patent/AR093346A2/es unknown
Patent Citations (2)
Publication number | Priority date | Publication date | Assignee | Title |
---|---|---|---|---|
WO1997030163A1 (en) * | 1996-02-14 | 1997-08-21 | The Governors Of The University Of Alberta | Plants having enhanced nitrogen assimilation/metabolism |
WO2001055433A2 (en) * | 2000-01-28 | 2001-08-02 | The Governors Of The University Of Alberta | Tissue-specific expression of target genes in plants |
Non-Patent Citations (4)
Title |
---|
Good AG, et al.Can less yield more? Is reducing nutrient input intotheenvironment compatible with maintaining crop production?.Trends in Plant Science9 12.2004,9(12),第597-605页. |
Good AG, et al.Can less yield more? Is reducing nutrient input intotheenvironment compatible with maintaining crop production?.Trends in Plant Science9 12.2004,9(12),第597-605页. * |
Iturriaga G, et al.Expression of desiccation-related proteins from theresurrectionplant Craterostigma plantagineum in transgenictobacco.Plant Molecular Biology20 3.1992,20(3),第555-558页. |
Iturriaga G, et al.Expression of desiccation-related proteins from theresurrectionplant Craterostigma plantagineum in transgenictobacco.Plant Molecular Biology20 3.1992,20(3),第555-558页. * |
Also Published As
Publication number | Publication date |
---|---|
CA2634925C (en) | 2015-06-23 |
EP1983819A4 (en) | 2009-09-02 |
AU2006330817B2 (en) | 2012-07-19 |
AR059119A1 (es) | 2008-03-12 |
ZA200805007B (en) | 2009-08-26 |
WO2007076115A3 (en) | 2008-06-19 |
BRPI0620312A2 (pt) | 2011-11-08 |
US20070162995A1 (en) | 2007-07-12 |
CN101378651A (zh) | 2009-03-04 |
US8288611B2 (en) | 2012-10-16 |
CA2634925A1 (en) | 2007-07-05 |
AU2006330817A1 (en) | 2007-07-05 |
US8642840B2 (en) | 2014-02-04 |
WO2007076115A2 (en) | 2007-07-05 |
AR093346A2 (es) | 2015-06-03 |
EP1983819A2 (en) | 2008-10-29 |
US20130097738A1 (en) | 2013-04-18 |
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Granted publication date: 20120314 Termination date: 20201221 |