JP6591993B2 - グルコースシロップ生産用組成物 - Google Patents

グルコースシロップ生産用組成物 Download PDF

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Publication number
JP6591993B2
JP6591993B2 JP2016550206A JP2016550206A JP6591993B2 JP 6591993 B2 JP6591993 B2 JP 6591993B2 JP 2016550206 A JP2016550206 A JP 2016550206A JP 2016550206 A JP2016550206 A JP 2016550206A JP 6591993 B2 JP6591993 B2 JP 6591993B2
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Japan
Prior art keywords
seq
amylase
pullulanase
glucoamylase
mature polypeptide
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English (en)
Japanese (ja)
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JP2017505129A5 (cg-RX-API-DMAC7.html
JP2017505129A (ja
Inventor
志朗 福山
志朗 福山
圭一 綾部
圭一 綾部
Original Assignee
ノボザイムス アクティーゼルスカブ
ノボザイムス アクティーゼルスカブ
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Classifications

    • A—HUMAN NECESSITIES
    • A23—FOODS OR FOODSTUFFS; TREATMENT THEREOF, NOT COVERED BY OTHER CLASSES
    • A23L—FOODS, FOODSTUFFS OR NON-ALCOHOLIC BEVERAGES, NOT OTHERWISE PROVIDED FOR; PREPARATION OR TREATMENT THEREOF
    • A23L29/00—Foods or foodstuffs containing additives; Preparation or treatment thereof
    • A23L29/30—Foods or foodstuffs containing additives; Preparation or treatment thereof containing carbohydrate syrups; containing sugars; containing sugar alcohols, e.g. xylitol; containing starch hydrolysates, e.g. dextrin
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12P—FERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P19/00—Preparation of compounds containing saccharide radicals
    • C12P19/14—Preparation of compounds containing saccharide radicals produced by the action of a carbohydrase (EC 3.2.x), e.g. by alpha-amylase, e.g. by cellulase, hemicellulase
    • A—HUMAN NECESSITIES
    • A23—FOODS OR FOODSTUFFS; TREATMENT THEREOF, NOT COVERED BY OTHER CLASSES
    • A23L—FOODS, FOODSTUFFS OR NON-ALCOHOLIC BEVERAGES, NOT OTHERWISE PROVIDED FOR; PREPARATION OR TREATMENT THEREOF
    • A23L29/00—Foods or foodstuffs containing additives; Preparation or treatment thereof
    • A23L29/30—Foods or foodstuffs containing additives; Preparation or treatment thereof containing carbohydrate syrups; containing sugars; containing sugar alcohols, e.g. xylitol; containing starch hydrolysates, e.g. dextrin
    • A23L29/35—Degradation products of starch, e.g. hydrolysates, dextrins; Enzymatically modified starches
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405—Glucanases
    • C12N9/2408—Glucanases acting on alpha -1,4-glucosidic bonds
    • C12N9/2411—Amylases
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405—Glucanases
    • C12N9/2408—Glucanases acting on alpha -1,4-glucosidic bonds
    • C12N9/2411—Amylases
    • C12N9/2428—Glucan 1,4-alpha-glucosidase (3.2.1.3), i.e. glucoamylase
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405—Glucanases
    • C12N9/2451—Glucanases acting on alpha-1,6-glucosidic bonds
    • C12N9/2457—Pullulanase (3.2.1.41)
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12P—FERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P19/00—Preparation of compounds containing saccharide radicals
    • C12P19/02—Monosaccharides
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12Y—ENZYMES
    • C12Y302/00—Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01—Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01001—Alpha-amylase (3.2.1.1)
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12Y—ENZYMES
    • C12Y302/00—Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01—Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01003—Glucan 1,4-alpha-glucosidase (3.2.1.3), i.e. glucoamylase
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12Y—ENZYMES
    • C12Y302/00—Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01—Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01041—Pullulanase (3.2.1.41)
    • C—CHEMISTRY; METALLURGY
    • C13—SUGAR INDUSTRY
    • C13K—SACCHARIDES OBTAINED FROM NATURAL SOURCES OR BY HYDROLYSIS OF NATURALLY OCCURRING DISACCHARIDES, OLIGOSACCHARIDES OR POLYSACCHARIDES
    • C13K1/00—Glucose; Glucose-containing syrups
    • C13K1/06—Glucose; Glucose-containing syrups obtained by saccharification of starch or raw materials containing starch
    • Y—GENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y02—TECHNOLOGIES OR APPLICATIONS FOR MITIGATION OR ADAPTATION AGAINST CLIMATE CHANGE
    • Y02E—REDUCTION OF GREENHOUSE GAS [GHG] EMISSIONS, RELATED TO ENERGY GENERATION, TRANSMISSION OR DISTRIBUTION
    • Y02E50/00—Technologies for the production of fuel of non-fossil origin
    • Y02E50/10—Biofuels, e.g. bio-diesel

Landscapes

  • Chemical & Material Sciences (AREA)
  • Life Sciences & Earth Sciences (AREA)
  • Organic Chemistry (AREA)
  • Health & Medical Sciences (AREA)
  • Engineering & Computer Science (AREA)
  • Wood Science & Technology (AREA)
  • Zoology (AREA)
  • Bioinformatics & Cheminformatics (AREA)
  • Genetics & Genomics (AREA)
  • General Health & Medical Sciences (AREA)
  • Biochemistry (AREA)
  • General Engineering & Computer Science (AREA)
  • Microbiology (AREA)
  • Biotechnology (AREA)
  • Molecular Biology (AREA)
  • Biomedical Technology (AREA)
  • Medicinal Chemistry (AREA)
  • Chemical Kinetics & Catalysis (AREA)
  • General Chemical & Material Sciences (AREA)
  • Nutrition Science (AREA)
  • Food Science & Technology (AREA)
  • Polymers & Plastics (AREA)
  • Emergency Medicine (AREA)
  • Preparation Of Compounds By Using Micro-Organisms (AREA)
  • Enzymes And Modification Thereof (AREA)
  • Jellies, Jams, And Syrups (AREA)
  • Micro-Organisms Or Cultivation Processes Thereof (AREA)
JP2016550206A 2014-02-07 2015-02-06 グルコースシロップ生産用組成物 Active JP6591993B2 (ja)

Applications Claiming Priority (5)

Application Number Priority Date Filing Date Title
EP14154239.9 2014-02-07
EP14154239 2014-02-07
EP14195687 2014-12-01
EP14195687.0 2014-12-01
PCT/EP2015/052545 WO2015118123A1 (en) 2014-02-07 2015-02-06 Compositions for producing glucose syrups

Publications (3)

Publication Number Publication Date
JP2017505129A JP2017505129A (ja) 2017-02-16
JP2017505129A5 JP2017505129A5 (cg-RX-API-DMAC7.html) 2018-03-15
JP6591993B2 true JP6591993B2 (ja) 2019-10-16

Family

ID=52464388

Family Applications (1)

Application Number Title Priority Date Filing Date
JP2016550206A Active JP6591993B2 (ja) 2014-02-07 2015-02-06 グルコースシロップ生産用組成物

Country Status (10)

Country Link
US (3) US10337041B2 (cg-RX-API-DMAC7.html)
EP (2) EP3712240B1 (cg-RX-API-DMAC7.html)
JP (1) JP6591993B2 (cg-RX-API-DMAC7.html)
CN (1) CN105960176A (cg-RX-API-DMAC7.html)
BR (1) BR112016018075B1 (cg-RX-API-DMAC7.html)
CA (1) CA2937818C (cg-RX-API-DMAC7.html)
HU (1) HUE064397T2 (cg-RX-API-DMAC7.html)
MX (1) MX385596B (cg-RX-API-DMAC7.html)
RU (2) RU2019134734A (cg-RX-API-DMAC7.html)
WO (1) WO2015118123A1 (cg-RX-API-DMAC7.html)

Families Citing this family (4)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
FR3091640B1 (fr) 2019-01-11 2021-06-11 Fermentalg Procédé de purification de phycocyanines
FR3091703B1 (fr) 2019-01-11 2021-02-12 Fermentalg Procédé d’extraction de phycocyanines
FR3092586A1 (fr) 2019-02-08 2020-08-14 Fermentalg Procédé optimisé d’exploitation industrielle d’algues rouges unicellulaires
US20250311734A1 (en) 2022-05-14 2025-10-09 Novozymes A/S Compositions and Methods for Preventing, Treating, Suppressing and/or Eliminatting Phytopathogenic Infestations and Infections

Family Cites Families (46)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
JPS5534046A (en) 1978-09-01 1980-03-10 Cpc International Inc Novel glucoamyrase having excellent heat resistance and production
US4335208A (en) 1980-03-11 1982-06-15 Novo Industri A/S Saccharification of starch hydrolysates
NO840200L (no) 1983-01-28 1984-07-30 Cefus Corp Glukoamylase cdna.
US4587215A (en) 1984-06-25 1986-05-06 Uop Inc. Highly thermostable amyloglucosidase
JPS62126989A (ja) 1985-11-26 1987-06-09 Godo Shiyusei Kk コルテイシウム属担子菌の生産する酵素を用いた澱粉質の無蒸煮糖化法
DE3886221T3 (de) 1987-09-04 2000-12-21 Novo Nordisk A/S, Bagsvaerd VERFAHREN ZUR HERSTELLUNG VON PROTEINPRODUKTEN IN -i(ASPERGILLUS) UND PROMOTOREN ZUR VERWENDUNG IN -i(ASPERGILLUS).
US5162210A (en) 1990-06-29 1992-11-10 Iowa State University Research Foundation Process for enzymatic hydrolysis of starch to glucose
ES2137222T3 (es) 1992-12-28 1999-12-16 Genencor Int Polulanasa, microorganismos que la producen, procedimientos para la preparacion de la misma y utilizaciones.
KR100511499B1 (ko) 1995-02-03 2005-12-21 노보자임스 에이/에스 소정 특성을 가지는 알파-아밀라제 돌연변이체를 디자인하는 방법
JP4290763B2 (ja) 1996-04-30 2009-07-08 ノボザイムス アクティーゼルスカブ α―アミラーゼ変異体
CA2259958A1 (en) 1996-07-24 1998-01-29 Martin Allen Protein engineering of glucoamylase to increase ph optimum, substrate specificity and thermostability
KR100808499B1 (ko) 1997-11-26 2008-02-29 노보자임스 에이/에스 열안정성 글루코아밀라제
CA2331340A1 (en) 1998-07-15 2000-01-27 Novozymes A/S Glucoamylase variants
WO2001004273A2 (en) 1999-07-09 2001-01-18 Novozymes A/S Glucoamylase variant
AU2002213841A1 (en) * 2000-11-10 2002-05-21 Novozymes A/S Secondary liquefaction of starch in ethanol production
ES2278210T3 (es) 2002-12-17 2007-08-01 Novozymes A/S Alfa-amilasas termoestables.
JP2007526748A (ja) 2003-06-25 2007-09-20 ノボザイムス アクティーゼルスカブ 澱粉加工用酵素
US7618795B2 (en) * 2003-06-25 2009-11-17 Novozymes A/S Starch process
WO2004113551A1 (en) * 2003-06-25 2004-12-29 Novozymes A/S Process for the hydrolysis of starch
CA2558603A1 (en) * 2004-03-08 2005-10-20 Syngenta Participations Ag Self-processing plants and plant parts
CA2566252A1 (en) 2004-05-13 2005-12-01 Novozymes North America, Inc. A process of producing a fermentation product
DK1824970T3 (da) 2004-11-30 2011-10-31 Genencor Int Thricoderma reesei-glycoamylase og homologer heraf
EP1831384B1 (en) 2004-12-22 2015-08-12 Novozymes A/S Polypeptides having glucoamylase activity and polynucleotides encoding same
WO2007144393A1 (en) * 2006-06-15 2007-12-21 Novozymes, Inc. Mashing process
JP5568307B2 (ja) 2006-10-10 2014-08-06 ダニスコ・ユーエス・インク、ジェネンコー・ディビジョン 修飾された性質を有するグルコアミラーゼ変異体
CN101049133B (zh) * 2007-04-26 2011-07-06 华南理工大学 小麦b淀粉多酶法制取发酵用含氮糖浆的方法
ES2504983T3 (es) 2007-10-09 2014-10-09 Danisco Us Inc. Variantes de glucoamilasa
CN101918525A (zh) 2007-12-12 2010-12-15 诺维信公司 糖化方法
BRPI0819869B1 (pt) * 2007-12-12 2019-07-16 Novozymes A/S Processo enzimático para a produção de um mosto de cervejeiro a partir de cereal não maltado, e, uso de um processo para produção de cerveja
US20120164695A1 (en) 2009-08-19 2012-06-28 Danisco Us Inc. Combinatorial variants of glucoamylase with improved specific activity and/or thermostability
EP2467474A1 (en) 2009-08-19 2012-06-27 Danisco A/S Variants of glucoamylase
CN102869771B (zh) 2009-11-30 2017-05-24 诺维信公司 具有葡糖淀粉酶活性的多肽和编码该多肽的多核苷酸
CA2782036C (en) 2009-12-01 2019-01-15 Novozymes A/S Polypeptides having glucoamylase activity and polynucleotides encoding same
CA2795806C (en) 2010-04-14 2019-01-08 Novozymes A/S Polypeptides having glucoamylase activity and polynucleotides encoding same
CN103298932B (zh) 2010-11-08 2016-08-10 诺维信公司 具有葡糖淀粉酶活性的多肽和编码该多肽的多核苷酸
KR101188113B1 (ko) * 2010-12-23 2012-10-05 대상 주식회사 유동성이 향상된 쌀엿 제조방법
JP5928089B2 (ja) * 2011-04-04 2016-06-01 国立大学法人弘前大学 イソマルターゼ
US9534239B2 (en) 2011-09-06 2017-01-03 Novozymes A/S Glucoamylase variants and polynucleotides encoding same
EP2766490B1 (en) 2011-10-11 2017-04-12 Novozymes North America, Inc. Liquefaction process for producing fermentation products
FR2989892B1 (fr) * 2012-04-26 2014-05-16 Oreal Composition cosmetique comprenant un oligosaccharide phosphoryle et un polysaccharide
MX2015002099A (es) * 2012-08-22 2015-05-11 Dupont Nutrition Biosci Aps Variantes que tienen actividad glucoamilasa.
ES2835313T3 (es) * 2013-04-30 2021-06-22 Novozymes As Variantes de glucoamilasa y polinucleótidos que las codifican
US10450551B2 (en) * 2013-08-30 2019-10-22 Novozymes A/S Enzyme composition and uses thereof
CN105960457B (zh) * 2014-01-22 2021-09-03 诺维信公司 普鲁兰酶变体以及编码它们的多核苷酸
ES2743515T3 (es) * 2014-10-23 2020-02-19 Novozymes As Variantes de glucoamilasa y polinucleótidos que codifican las mismas
MX2017007102A (es) * 2014-12-01 2017-08-24 Novozymes As Produccion mejorada de jarabes de glucosa.

Also Published As

Publication number Publication date
US11136607B2 (en) 2021-10-05
US11667938B2 (en) 2023-06-06
US20190256878A1 (en) 2019-08-22
RU2706297C2 (ru) 2019-11-15
CA2937818C (en) 2023-08-01
HUE064397T2 (hu) 2024-03-28
RU2016135029A3 (cg-RX-API-DMAC7.html) 2018-10-16
EP3102051B1 (en) 2020-04-08
US20170159090A1 (en) 2017-06-08
CA2937818A1 (en) 2015-08-13
EP3102051A1 (en) 2016-12-14
US10337041B2 (en) 2019-07-02
RU2019134734A (ru) 2019-11-18
MX385596B (es) 2025-03-18
JP2017505129A (ja) 2017-02-16
EP3712240A1 (en) 2020-09-23
RU2016135029A (ru) 2018-03-13
US20210395791A1 (en) 2021-12-23
WO2015118123A1 (en) 2015-08-13
EP3712240B1 (en) 2023-09-27
BR112016018075A2 (pt) 2018-02-20
CN105960176A (zh) 2016-09-21
MX2016010031A (es) 2016-11-15
BR112016018075B1 (pt) 2022-01-18

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