JP5677839B2 - 突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物 - Google Patents

突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物 Download PDF

Info

Publication number
JP5677839B2
JP5677839B2 JP2010515401A JP2010515401A JP5677839B2 JP 5677839 B2 JP5677839 B2 JP 5677839B2 JP 2010515401 A JP2010515401 A JP 2010515401A JP 2010515401 A JP2010515401 A JP 2010515401A JP 5677839 B2 JP5677839 B2 JP 5677839B2
Authority
JP
Japan
Prior art keywords
fatb
seq
nucleic acid
mutant
plant
Prior art date
Legal status (The legal status is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the status listed.)
Expired - Fee Related
Application number
JP2010515401A
Other languages
English (en)
Japanese (ja)
Other versions
JP2010532667A (ja
JP2010532667A5 (enExample
Inventor
ラハ,ベンヤミン
デン・ブール,バルト
ランベルト,バルト
Current Assignee (The listed assignees may be inaccurate. Google has not performed a legal analysis and makes no representation or warranty as to the accuracy of the list.)
Bayer CropScience NV
Original Assignee
Bayer CropScience NV
Priority date (The priority date is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the date listed.)
Filing date
Publication date
Application filed by Bayer CropScience NV filed Critical Bayer CropScience NV
Publication of JP2010532667A publication Critical patent/JP2010532667A/ja
Publication of JP2010532667A5 publication Critical patent/JP2010532667A5/ja
Application granted granted Critical
Publication of JP5677839B2 publication Critical patent/JP5677839B2/ja
Expired - Fee Related legal-status Critical Current
Anticipated expiration legal-status Critical

Links

Images

Classifications

    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12QMEASURING OR TESTING PROCESSES INVOLVING ENZYMES, NUCLEIC ACIDS OR MICROORGANISMS; COMPOSITIONS OR TEST PAPERS THEREFOR; PROCESSES OF PREPARING SUCH COMPOSITIONS; CONDITION-RESPONSIVE CONTROL IN MICROBIOLOGICAL OR ENZYMOLOGICAL PROCESSES
    • C12Q1/00Measuring or testing processes involving enzymes, nucleic acids or microorganisms; Compositions therefor; Processes of preparing such compositions
    • C12Q1/68Measuring or testing processes involving enzymes, nucleic acids or microorganisms; Compositions therefor; Processes of preparing such compositions involving nucleic acids
    • C12Q1/6876Nucleic acid products used in the analysis of nucleic acids, e.g. primers or probes
    • C12Q1/6888Nucleic acid products used in the analysis of nucleic acids, e.g. primers or probes for detection or identification of organisms
    • C12Q1/6895Nucleic acid products used in the analysis of nucleic acids, e.g. primers or probes for detection or identification of organisms for plants, fungi or algae
    • AHUMAN NECESSITIES
    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01HNEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
    • A01H1/00Processes for modifying genotypes ; Plants characterised by associated natural traits
    • A01H1/04Processes of selection involving genotypic or phenotypic markers; Methods of using phenotypic markers for selection
    • A01H1/045Processes of selection involving genotypic or phenotypic markers; Methods of using phenotypic markers for selection using molecular markers
    • AHUMAN NECESSITIES
    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01HNEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
    • A01H5/00Angiosperms, i.e. flowering plants, characterised by their plant parts; Angiosperms characterised otherwise than by their botanic taxonomy
    • A01H5/10Seeds
    • AHUMAN NECESSITIES
    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01HNEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
    • A01H6/00Angiosperms, i.e. flowering plants, characterised by their botanic taxonomy
    • A01H6/20Brassicaceae, e.g. canola, broccoli or rucola
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12NMICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N15/00Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
    • C12N15/09Recombinant DNA-technology
    • C12N15/63Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
    • C12N15/79Vectors or expression systems specially adapted for eukaryotic hosts
    • C12N15/82Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
    • C12N15/8241Phenotypically and genetically modified plants via recombinant DNA technology
    • C12N15/8242Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits
    • C12N15/8243Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits involving biosynthetic or metabolic pathways, i.e. metabolic engineering, e.g. nicotine, caffeine
    • C12N15/8247Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits involving biosynthetic or metabolic pathways, i.e. metabolic engineering, e.g. nicotine, caffeine involving modified lipid metabolism, e.g. seed oil composition
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12NMICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14Hydrolases (3)
    • C12N9/16Hydrolases (3) acting on ester bonds (3.1)
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12QMEASURING OR TESTING PROCESSES INVOLVING ENZYMES, NUCLEIC ACIDS OR MICROORGANISMS; COMPOSITIONS OR TEST PAPERS THEREFOR; PROCESSES OF PREPARING SUCH COMPOSITIONS; CONDITION-RESPONSIVE CONTROL IN MICROBIOLOGICAL OR ENZYMOLOGICAL PROCESSES
    • C12Q2600/00Oligonucleotides characterized by their use
    • C12Q2600/156Polymorphic or mutational markers
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12QMEASURING OR TESTING PROCESSES INVOLVING ENZYMES, NUCLEIC ACIDS OR MICROORGANISMS; COMPOSITIONS OR TEST PAPERS THEREFOR; PROCESSES OF PREPARING SUCH COMPOSITIONS; CONDITION-RESPONSIVE CONTROL IN MICROBIOLOGICAL OR ENZYMOLOGICAL PROCESSES
    • C12Q2600/00Oligonucleotides characterized by their use
    • C12Q2600/158Expression markers
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12QMEASURING OR TESTING PROCESSES INVOLVING ENZYMES, NUCLEIC ACIDS OR MICROORGANISMS; COMPOSITIONS OR TEST PAPERS THEREFOR; PROCESSES OF PREPARING SUCH COMPOSITIONS; CONDITION-RESPONSIVE CONTROL IN MICROBIOLOGICAL OR ENZYMOLOGICAL PROCESSES
    • C12Q2600/00Oligonucleotides characterized by their use
    • C12Q2600/172Haplotypes

Landscapes

  • Life Sciences & Earth Sciences (AREA)
  • Health & Medical Sciences (AREA)
  • Chemical & Material Sciences (AREA)
  • Genetics & Genomics (AREA)
  • Engineering & Computer Science (AREA)
  • Organic Chemistry (AREA)
  • Zoology (AREA)
  • Wood Science & Technology (AREA)
  • Biotechnology (AREA)
  • Bioinformatics & Cheminformatics (AREA)
  • General Health & Medical Sciences (AREA)
  • Botany (AREA)
  • General Engineering & Computer Science (AREA)
  • Molecular Biology (AREA)
  • Analytical Chemistry (AREA)
  • Proteomics, Peptides & Aminoacids (AREA)
  • Biochemistry (AREA)
  • Microbiology (AREA)
  • Developmental Biology & Embryology (AREA)
  • Biomedical Technology (AREA)
  • Environmental Sciences (AREA)
  • Physics & Mathematics (AREA)
  • Biophysics (AREA)
  • Physiology (AREA)
  • Mycology (AREA)
  • Immunology (AREA)
  • Medicinal Chemistry (AREA)
  • Cell Biology (AREA)
  • Plant Pathology (AREA)
  • Nutrition Science (AREA)
  • Oil, Petroleum & Natural Gas (AREA)
  • Spectroscopy & Molecular Physics (AREA)
  • Natural Medicines & Medicinal Plants (AREA)
  • Measuring Or Testing Involving Enzymes Or Micro-Organisms (AREA)
  • Breeding Of Plants And Reproduction By Means Of Culturing (AREA)
  • Enzymes And Modification Thereof (AREA)
  • Micro-Organisms Or Cultivation Processes Thereof (AREA)
  • Coloring Foods And Improving Nutritive Qualities (AREA)
  • Preparation Of Compounds By Using Micro-Organisms (AREA)
JP2010515401A 2007-07-09 2008-07-07 突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物 Expired - Fee Related JP5677839B2 (ja)

Applications Claiming Priority (5)

Application Number Priority Date Filing Date Title
EP07075568.1 2007-07-09
EP07075568 2007-07-09
US95894507P 2007-07-10 2007-07-10
US60/958,945 2007-07-10
PCT/EP2008/005551 WO2009007091A2 (en) 2007-07-09 2008-07-07 Brassica plant comprising mutant fatty acyl-acp thioesterase alleles

Publications (3)

Publication Number Publication Date
JP2010532667A JP2010532667A (ja) 2010-10-14
JP2010532667A5 JP2010532667A5 (enExample) 2011-08-25
JP5677839B2 true JP5677839B2 (ja) 2015-02-25

Family

ID=40076671

Family Applications (1)

Application Number Title Priority Date Filing Date
JP2010515401A Expired - Fee Related JP5677839B2 (ja) 2007-07-09 2008-07-07 突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物

Country Status (15)

Country Link
US (2) US8981180B2 (enExample)
EP (1) EP2167667B1 (enExample)
JP (1) JP5677839B2 (enExample)
CN (1) CN101688220B (enExample)
AR (1) AR067480A1 (enExample)
AU (1) AU2008274551B2 (enExample)
CA (2) CA2692687C (enExample)
DK (1) DK2167667T3 (enExample)
EA (1) EA030862B1 (enExample)
ES (1) ES2545083T3 (enExample)
PL (1) PL2167667T3 (enExample)
SI (1) SI2167667T1 (enExample)
UA (1) UA99471C2 (enExample)
WO (1) WO2009007091A2 (enExample)
ZA (2) ZA200908886B (enExample)

Families Citing this family (41)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
ES2342466T7 (es) 2003-11-21 2012-11-19 Givaudan Sa P-mentanocarboxamidas n-substituidas.
JP5827233B2 (ja) 2009-10-22 2015-12-02 ダウ アグロサイエンシィズ エルエルシー 脂肪酸生合成に関与する植物遺伝子をターゲッティングする改変ジンクフィンガータンパク質
WO2011060946A1 (en) 2009-11-20 2011-05-26 Bayer Bioscience N.V. Brassica plants comprising mutant fad3 alleles
EP3053432A1 (en) * 2009-12-18 2016-08-10 Cargill, Incorporated Brassica plants yielding oils with a low total saturated fatty acid content
CN102811617A (zh) 2010-01-22 2012-12-05 拜耳知识产权有限责任公司 杀螨和/或杀虫活性物质结合物
CN102892903A (zh) * 2010-05-12 2013-01-23 农业基因遗传学有限公司 褐色中脉3基因特异性标志物在玉米中用于性状基因渗入的用途
US9695434B2 (en) * 2010-05-25 2017-07-04 Cargill, Incorporated Brassica plants yielding oils with a low alpha linolenic acid content
AU2011258362B2 (en) * 2010-05-25 2016-07-07 Pioneer Hi-Bred International, Inc. Brassica plants yielding oils with a low alpha linolenic acid content
WO2012061647A2 (en) * 2010-11-03 2012-05-10 Solazyme, Inc. Microbial oils with lowered pour points, dielectric fluids produced therefrom, and related methods
JP5860471B2 (ja) 2010-11-15 2016-02-16 バイエル・インテレクチュアル・プロパティ・ゲゼルシャフト・ミット・ベシュレンクテル・ハフツングBayer Intellectual Property GmbH N−アリールピラゾール(チオ)カルボキサミド類
WO2013020985A1 (en) 2011-08-10 2013-02-14 Bayer Intellectual Property Gmbh Active compound combinations comprising specific tetramic acid derivatives
US9968041B2 (en) 2012-07-06 2018-05-15 Bayer Cropscience Nv Soybean ROD1 gene sequences and uses thereof
CA2878289A1 (en) 2012-07-06 2014-01-09 Bayer Cropscience Nv Brassica plants with modified seed oil composition
WO2014006158A1 (en) 2012-07-06 2014-01-09 Bayer Cropscience Nv Brassica rod1 gene sequences and uses thereof
US9428459B2 (en) 2012-12-19 2016-08-30 Bayer Cropscience Ag Difluoromethyl-nicotinic- tetrahydronaphtyl carboxamides
US9816079B2 (en) 2013-01-29 2017-11-14 Terravia Holdings, Inc. Variant thioesterases and methods of use
US9567615B2 (en) 2013-01-29 2017-02-14 Terravia Holdings, Inc. Variant thioesterases and methods of use
EP2971024A4 (en) * 2013-03-15 2016-11-16 Terravia Holdings Inc THIOESTERASES AND CELLS FOR THE MANUFACTURE OF TAILOR-MADE OILS
US9783836B2 (en) 2013-03-15 2017-10-10 Terravia Holdings, Inc. Thioesterases and cells for production of tailored oils
US9290749B2 (en) 2013-03-15 2016-03-22 Solazyme, Inc. Thioesterases and cells for production of tailored oils
WO2016014968A1 (en) 2014-07-24 2016-01-28 Solazyme, Inc. Variant thioesterases and methods of use
BR112017005370A2 (pt) 2014-09-18 2017-12-12 Terravia Holdings Inc acil-acp tioesterases e mutantes das mesmas
CN107624135A (zh) 2015-04-28 2018-01-23 拜尔作物科学公司 具有修饰的种子油组成的芸苔属植物
EP3464600A1 (en) * 2016-05-25 2019-04-10 Cargill, Incorporated Engineered nucleases to generate deletion mutants in plants
US11517025B2 (en) 2016-08-12 2022-12-06 Cargill, Incorporated Speciality low saturates canola oil
US10440915B2 (en) 2016-09-01 2019-10-15 Cargill, Incorporated Canola cultivar 15RH0611
US10440916B2 (en) 2016-09-06 2019-10-15 Cargill, Incorporated Canola cultivar 15RH0612
US10440909B2 (en) 2016-09-06 2019-10-15 Cargill, Incorporated Canola cultivar 15RH0613
TWI873082B (zh) * 2017-08-17 2025-02-21 日商莎卡達種子股份有限公司 甘藍(Brassica oleracea)植物之異型檢出法
US20220033835A1 (en) * 2018-09-14 2022-02-03 Chuanxin Sun Rice plant material resistant against biotic stress
AU2020204657A1 (en) 2019-01-04 2021-07-29 Pioneer Hi-Bred International, Inc. Engineered nucleases to generate mutations in plants
CA3047768A1 (en) 2019-06-21 2020-12-21 BASF Agricultural Solutions Seed US LLC Canola hybrid variety 7cn0425
CA3165554A1 (en) * 2020-01-31 2021-08-05 Qiwei SHAN Increased of saturated fat in soybean
US11997966B2 (en) 2020-04-24 2024-06-04 BASF Agricultural Solutions Seed US LLC Canola hybrid variety 7CN0020
US11997965B2 (en) 2020-04-24 2024-06-04 BASF Agricultural Solutions Seed US LLC Canola hybrid variety 8CN0001
US11672218B2 (en) 2020-04-24 2023-06-13 BASF Agricultural Solutions Seed US LLC Canola hybrid variety 7CN0065
CA3201992A1 (en) 2020-12-21 2022-06-30 Thi Ninh Thuan NGUYEN Brassica napus plants comprising an improved fertility restorer
JP2025507908A (ja) 2022-03-01 2025-03-21 ビーエーエスエフ アグリカルチュラル ソリューションズ シード ユーエス エルエルシー Cas12aニッカーゼ
CN114958880B (zh) * 2022-04-06 2023-08-29 吉林大学 大豆脂酰-酰基载体蛋白硫酯酶GmFATA2基因及应用
WO2025045884A1 (en) 2023-08-31 2025-03-06 Basf Agricultural Solutions Us Llc Class 2 type v crispr-cas prime editing
WO2025083165A1 (en) 2023-10-19 2025-04-24 Basf Agricultural Solutions Us Llc Brassica napus plants having enhanced blackleg resistance

Family Cites Families (16)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
DK0476093T3 (da) * 1990-04-04 1997-06-02 Pioneer Hi Bred Int Fremstilling af forbedret rapsfrø, der udviser et reduceret indhold af mættet fedtsyre
US5344771A (en) 1990-04-26 1994-09-06 Calgene, Inc. Plant thiosterases
US5512482A (en) 1990-04-26 1996-04-30 Calgene, Inc. Plant thioesterases
DE69113635T3 (de) 1990-12-20 2000-06-08 E.I. Du Pont De Nemours And Co., Wilmington Nukleotidsequenzen der sojabohnen-acyl-acp-thioesterasegene.
US5945585A (en) 1990-12-20 1999-08-31 E. I. Du Pont De Nemours And Company Specific for palmitoyl, stearoyl and oleoyl-alp thioesters nucleic acid fragments encoding acyl-acp thiosesterase enzymes and the use of these fragments in altering plant oil composition
US5455167A (en) 1991-05-21 1995-10-03 Calgene Inc. Medium-chain thioesterases in plants
PT969102E (pt) 1991-09-24 2008-03-25 Keygene Nv Iniciadores, grupos e conjuntos de fragmentos de restrição usados na amplificação selectiva de fragmentos de restrição
US5850022A (en) 1992-10-30 1998-12-15 Calgene, Inc. Production of myristate in plant cells
WO1995013390A2 (en) 1993-11-10 1995-05-18 Calgene, Inc. Plant acyl acp thioesterase sequences
CA2198222C (en) * 1994-08-31 2011-10-25 William Dean Hitz Nucleotide sequences of canola and soybean palmitoyl-acp thioesterase genes and their use in the regulation of fatty acid content of the oils of soybean and canola plants
US5625130A (en) * 1995-03-07 1997-04-29 Pioneer Hi-Bred International, Inc. Oilseed Brassica bearing an endogenous oil wherein the levels of oleic, alpha-linolenic, and saturated fatty acids are simultaneously provided in an atypical highly beneficial distribution via genetic control
US5985557A (en) 1996-01-24 1999-11-16 Third Wave Technologies, Inc. Invasive cleavage of nucleic acids
US7067722B2 (en) * 1999-08-26 2006-06-27 Monsanto Technology Llc Nucleic acid sequences and methods of use for the production of plants with modified polyunsaturated fatty acids
BR0112716A (pt) 2000-07-25 2003-06-24 Calgene Llc Sequencias de ácidos nucléicos codificando beta-cetoacil-acp sintase e aplicações das mesmas
AU2003277812A1 (en) * 2002-06-21 2004-01-06 Monsanto Technology Llc Thioesterase-related nucleic acid sequences and methods
ES2388066T3 (es) * 2004-10-08 2012-10-08 Dow Agrosciences Llc Plantas con niveles nulos o reducidos de ácidos grasos saturados en sus semillas y aceite derivado de dichas semillas

Also Published As

Publication number Publication date
JP2010532667A (ja) 2010-10-14
WO2009007091A2 (en) 2009-01-15
CN101688220A (zh) 2010-03-31
EP2167667A2 (en) 2010-03-31
US20150232951A1 (en) 2015-08-20
AU2008274551B2 (en) 2014-05-29
US10837066B2 (en) 2020-11-17
EA201070114A1 (ru) 2010-08-30
EP2167667B1 (en) 2015-05-20
DK2167667T3 (en) 2015-07-27
AU2008274551A1 (en) 2009-01-15
UA99471C2 (ru) 2012-08-27
ES2545083T3 (es) 2015-09-08
ZA201009002B (en) 2011-07-27
ZA200908886B (en) 2011-02-23
WO2009007091A3 (en) 2009-07-23
EA030862B1 (ru) 2018-10-31
US8981180B2 (en) 2015-03-17
AR067480A1 (es) 2009-10-14
CN101688220B (zh) 2017-04-05
CA2692687A1 (en) 2009-01-15
CA2692687C (en) 2021-10-26
SI2167667T1 (sl) 2016-03-31
PL2167667T3 (pl) 2015-10-30
CA3128584A1 (en) 2009-01-15
US20110145944A1 (en) 2011-06-16

Similar Documents

Publication Publication Date Title
JP5677839B2 (ja) 突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物
AU2010321249B2 (en) Brassica plants comprising mutant FAD3 alleles
Richmond et al. A defect in β-oxidation causes abnormal inflorescence development in Arabidopsis
JP2018085986A (ja) オメガ9品質カラシナ
US10494642B2 (en) Brassica plants with modified seed oil composition
NZ788009A (en) Elite event canola ns-b50027-4
KR20220068223A (ko) 브라시카 윤케아 계통 nubj1207
AU2008200860A1 (en) Brassica juncea lines with high oleic acid profile in seed oil
HK1139981A (en) Brassica plant comprising mutant fatty acyl-acp thioesterase alleles
WO2016176476A1 (en) Brassica plants with modified seed oil composition

Legal Events

Date Code Title Description
A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20110707

A621 Written request for application examination

Free format text: JAPANESE INTERMEDIATE CODE: A621

Effective date: 20110707

A711 Notification of change in applicant

Free format text: JAPANESE INTERMEDIATE CODE: A712

Effective date: 20120613

A131 Notification of reasons for refusal

Free format text: JAPANESE INTERMEDIATE CODE: A131

Effective date: 20130806

A02 Decision of refusal

Free format text: JAPANESE INTERMEDIATE CODE: A02

Effective date: 20140617

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20141017

A911 Transfer to examiner for re-examination before appeal (zenchi)

Free format text: JAPANESE INTERMEDIATE CODE: A911

Effective date: 20141027

TRDD Decision of grant or rejection written
A01 Written decision to grant a patent or to grant a registration (utility model)

Free format text: JAPANESE INTERMEDIATE CODE: A01

Effective date: 20141216

A61 First payment of annual fees (during grant procedure)

Free format text: JAPANESE INTERMEDIATE CODE: A61

Effective date: 20150105

R150 Certificate of patent or registration of utility model

Ref document number: 5677839

Country of ref document: JP

Free format text: JAPANESE INTERMEDIATE CODE: R150

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

S111 Request for change of ownership or part of ownership

Free format text: JAPANESE INTERMEDIATE CODE: R313113

R350 Written notification of registration of transfer

Free format text: JAPANESE INTERMEDIATE CODE: R350

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

R250 Receipt of annual fees

Free format text: JAPANESE INTERMEDIATE CODE: R250

LAPS Cancellation because of no payment of annual fees