JP7549444B2 - 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品 - Google Patents

微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品 Download PDF

Info

Publication number
JP7549444B2
JP7549444B2 JP2018569171A JP2018569171A JP7549444B2 JP 7549444 B2 JP7549444 B2 JP 7549444B2 JP 2018569171 A JP2018569171 A JP 2018569171A JP 2018569171 A JP2018569171 A JP 2018569171A JP 7549444 B2 JP7549444 B2 JP 7549444B2
Authority
JP
Japan
Prior art keywords
cells
microalgae
culture
heterotrophic
growth
Prior art date
Legal status (The legal status is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the status listed.)
Active
Application number
JP2018569171A
Other languages
English (en)
Japanese (ja)
Other versions
JP2019527052A (ja
JP2019527052A5 (https=
Inventor
シュール,ロバート,ジェー.
クーンル,アデルハイト,アール.
Original Assignee
クーンル アグロシステムズ,インコーポレイテッド
Priority date (The priority date is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the date listed.)
Filing date
Publication date
Application filed by クーンル アグロシステムズ,インコーポレイテッド filed Critical クーンル アグロシステムズ,インコーポレイテッド
Publication of JP2019527052A publication Critical patent/JP2019527052A/ja
Publication of JP2019527052A5 publication Critical patent/JP2019527052A5/ja
Priority to JP2022133647A priority Critical patent/JP2022177001A/ja
Application granted granted Critical
Publication of JP7549444B2 publication Critical patent/JP7549444B2/ja
Priority to JP2024225562A priority patent/JP2025041773A/ja
Active legal-status Critical Current
Anticipated expiration legal-status Critical

Links

Classifications

    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12NMICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N15/00Mutation or genetic engineering; DNA or RNA concerning genetic engineering, vectors, e.g. plasmids, or their isolation, preparation or purification; Use of hosts therefor
    • C12N15/09Recombinant DNA-technology
    • C12N15/63Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
    • C12N15/79Vectors or expression systems specially adapted for eukaryotic hosts
    • C12N15/82Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
    • AHUMAN NECESSITIES
    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01NPRESERVATION OF BODIES OF HUMANS OR ANIMALS OR PLANTS OR PARTS THEREOF; BIOCIDES, e.g. AS DISINFECTANTS, AS PESTICIDES OR AS HERBICIDES; PEST REPELLANTS OR ATTRACTANTS; PLANT GROWTH REGULATORS
    • A01N65/00Biocides, pest repellants or attractants, or plant growth regulators containing material from algae, lichens, bryophyta, multi-cellular fungi or plants, or extracts thereof
    • A01N65/03Algae
    • AHUMAN NECESSITIES
    • A23FOODS OR FOODSTUFFS; TREATMENT THEREOF, NOT COVERED BY OTHER CLASSES
    • A23KFODDER
    • A23K20/00Accessory food factors for animal feeding-stuffs
    • A23K20/10Organic substances
    • AHUMAN NECESSITIES
    • A23FOODS OR FOODSTUFFS; TREATMENT THEREOF, NOT COVERED BY OTHER CLASSES
    • A23KFODDER
    • A23K50/00Feeding-stuffs specially adapted for particular animals
    • A23K50/80Feeding-stuffs specially adapted for particular animals for aquatic animals, e.g. fish, crustaceans or molluscs
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12NMICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N1/00Microorganisms; Compositions thereof; Processes of propagating, maintaining or preserving microorganisms or compositions thereof; Processes of preparing or isolating a composition containing a microorganism; Culture media therefor
    • C12N1/12Unicellular algae; Culture media therefor
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12PFERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P23/00Preparation of compounds containing a cyclohexene ring having an unsaturated side chain containing at least ten carbon atoms bound by conjugated double bonds, e.g. carotenes
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12PFERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P3/00Preparation of elements or inorganic compounds except carbon dioxide
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12PFERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P5/00Preparation of hydrocarbons or halogenated hydrocarbons
    • C12P5/02Preparation of hydrocarbons or halogenated hydrocarbons acyclic
    • C12P5/026Unsaturated compounds, i.e. alkenes, alkynes or allenes
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12PFERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P7/00Preparation of oxygen-containing organic compounds
    • C12P7/24Preparation of oxygen-containing organic compounds containing a carbonyl group
    • C12P7/26Ketones
    • CCHEMISTRY; METALLURGY
    • C12BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12PFERMENTATION OR ENZYME-USING PROCESSES TO SYNTHESISE A DESIRED CHEMICAL COMPOUND OR COMPOSITION OR TO SEPARATE OPTICAL ISOMERS FROM A RACEMIC MIXTURE
    • C12P9/00Preparation of organic compounds containing a metal or atom other than H, N, C, O, S or halogen
    • GPHYSICS
    • G01MEASURING; TESTING
    • G01NINVESTIGATING OR ANALYSING MATERIALS BY DETERMINING THEIR CHEMICAL OR PHYSICAL PROPERTIES
    • G01N33/00Investigating or analysing materials by specific methods not covered by groups G01N1/00 - G01N31/00
    • G01N33/48Biological material, e.g. blood, urine; Haemocytometers
    • G01N33/50Chemical analysis of biological material, e.g. blood, urine; Testing involving biospecific ligand binding methods; Immunological testing
    • G01N33/5097Chemical analysis of biological material, e.g. blood, urine; Testing involving biospecific ligand binding methods; Immunological testing involving plant cells
    • GPHYSICS
    • G01MEASURING; TESTING
    • G01NINVESTIGATING OR ANALYSING MATERIALS BY DETERMINING THEIR CHEMICAL OR PHYSICAL PROPERTIES
    • G01N2333/00Assays involving biological materials from specific organisms or of a specific nature
    • G01N2333/405Assays involving biological materials from specific organisms or of a specific nature from algae
    • YGENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y02TECHNOLOGIES OR APPLICATIONS FOR MITIGATION OR ADAPTATION AGAINST CLIMATE CHANGE
    • Y02EREDUCTION OF GREENHOUSE GAS [GHG] EMISSIONS, RELATED TO ENERGY GENERATION, TRANSMISSION OR DISTRIBUTION
    • Y02E50/00Technologies for the production of fuel of non-fossil origin
    • Y02E50/30Fuel from waste, e.g. synthetic alcohol or diesel

Landscapes

  • Life Sciences & Earth Sciences (AREA)
  • Chemical & Material Sciences (AREA)
  • Engineering & Computer Science (AREA)
  • Health & Medical Sciences (AREA)
  • Organic Chemistry (AREA)
  • Zoology (AREA)
  • Wood Science & Technology (AREA)
  • Biotechnology (AREA)
  • Genetics & Genomics (AREA)
  • General Health & Medical Sciences (AREA)
  • Bioinformatics & Cheminformatics (AREA)
  • Microbiology (AREA)
  • Biochemistry (AREA)
  • General Engineering & Computer Science (AREA)
  • General Chemical & Material Sciences (AREA)
  • Chemical Kinetics & Catalysis (AREA)
  • Biomedical Technology (AREA)
  • Polymers & Plastics (AREA)
  • Cell Biology (AREA)
  • Molecular Biology (AREA)
  • Immunology (AREA)
  • Food Science & Technology (AREA)
  • Urology & Nephrology (AREA)
  • Physics & Mathematics (AREA)
  • Medicinal Chemistry (AREA)
  • Plant Pathology (AREA)
  • Hematology (AREA)
  • Botany (AREA)
  • Animal Husbandry (AREA)
  • Birds (AREA)
  • Pathology (AREA)
  • Biophysics (AREA)
  • Natural Medicines & Medicinal Plants (AREA)
  • Environmental Sciences (AREA)
  • Marine Sciences & Fisheries (AREA)
  • Dentistry (AREA)
  • Insects & Arthropods (AREA)
  • Mycology (AREA)
  • Analytical Chemistry (AREA)
  • General Physics & Mathematics (AREA)
JP2018569171A 2016-06-30 2017-06-30 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品 Active JP7549444B2 (ja)

Priority Applications (2)

Application Number Priority Date Filing Date Title
JP2022133647A JP2022177001A (ja) 2016-06-30 2022-08-24 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品
JP2024225562A JP2025041773A (ja) 2016-06-30 2024-12-20 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品

Applications Claiming Priority (3)

Application Number Priority Date Filing Date Title
US201662356896P 2016-06-30 2016-06-30
US62/356,896 2016-06-30
PCT/US2017/040486 WO2018006068A1 (en) 2016-06-30 2017-06-30 Improved heterotrophic production methods for microbial biomass and bioproducts

Related Child Applications (1)

Application Number Title Priority Date Filing Date
JP2022133647A Division JP2022177001A (ja) 2016-06-30 2022-08-24 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品

Publications (3)

Publication Number Publication Date
JP2019527052A JP2019527052A (ja) 2019-09-26
JP2019527052A5 JP2019527052A5 (https=) 2020-08-13
JP7549444B2 true JP7549444B2 (ja) 2024-09-11

Family

ID=59351112

Family Applications (3)

Application Number Title Priority Date Filing Date
JP2018569171A Active JP7549444B2 (ja) 2016-06-30 2017-06-30 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品
JP2022133647A Withdrawn JP2022177001A (ja) 2016-06-30 2022-08-24 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品
JP2024225562A Pending JP2025041773A (ja) 2016-06-30 2024-12-20 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品

Family Applications After (2)

Application Number Title Priority Date Filing Date
JP2022133647A Withdrawn JP2022177001A (ja) 2016-06-30 2022-08-24 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品
JP2024225562A Pending JP2025041773A (ja) 2016-06-30 2024-12-20 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品

Country Status (5)

Country Link
US (2) US11034968B2 (https=)
EP (1) EP3478850A1 (https=)
JP (3) JP7549444B2 (https=)
CA (1) CA3028227A1 (https=)
WO (1) WO2018006068A1 (https=)

Families Citing this family (16)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
US11419350B2 (en) 2016-07-01 2022-08-23 Corbion Biotech, Inc. Feed ingredients comprising lysed microbial cells
US11377675B2 (en) * 2019-01-16 2022-07-05 Kuehnle Agrosystems, Inc. Subterranean microalgae for production of microbial biomass, substances, and compositions
WO2021057709A1 (zh) * 2019-09-23 2021-04-01 山东拜昂生物技术有限公司 一种培养雨生红球藻生产虾青素的方法
US20220356504A1 (en) * 2019-09-23 2022-11-10 Bioalgo (Wf) Co., Ltd. Method for producing astaxanthin by heterotrophic culture of haematococcus pluvialis
CN110804639B (zh) * 2019-11-27 2022-04-08 中国科学院天津工业生物技术研究所 促进微藻虾青素酯化的方法
KR102445369B1 (ko) * 2019-12-03 2022-09-21 한양대학교 산학협력단 미세조류로부터 기능성 오일을 추출하는 방법 및 기능성 오일을 포함하는 유화액
EP4210663B1 (en) * 2020-09-10 2026-04-29 The Procter & Gamble Company Late-stage product differentiation process for personal care products
USPP34488P3 (en) 2020-10-30 2022-08-09 Kuehnle Agrosystems, Inc. Haematococcus microalga named ‘KAS1601-WG7’
KR20240014474A (ko) * 2021-05-27 2024-02-01 알.피.쉐러 테크놀러지즈 엘엘씨 포르밀글리신 함유 폴리펩티드의 절단을 조절하는 방법
EP4122330A1 (en) * 2021-07-22 2023-01-25 Alver World SA Methods of growing microalgae & products thereof
WO2023074226A1 (ja) * 2021-10-27 2023-05-04 本田技研工業株式会社 緑藻クラミドモナスの培養方法
CN115024428B (zh) * 2022-06-15 2024-05-24 广州希望饲料有限公司 一种具有改善水禽免疫力与肠道的无抗饲料添加剂及其制备方法
US20260047991A1 (en) * 2022-08-08 2026-02-19 University Of Florida Research Foundation, Inc. Stimuli-responsive chromogenic materials, methods of making and methods of use
WO2024258936A2 (en) * 2023-06-13 2024-12-19 Kuehnle Agrosystems, Inc. Improved edible compositions comprising an improved chlamydomonas reinhardtii component
KR102897913B1 (ko) * 2023-12-26 2025-12-12 전제은 미세조류로부터 관심 생성물을 고수율로 수득하기 위한 방법
CN120183511A (zh) * 2025-05-22 2025-06-20 成都英德生物医药设备有限公司 生物反应器的动态优化方法以及系统

Citations (5)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
US6022701A (en) 1996-02-01 2000-02-08 Ben-Gurion University Of The Negev Research And Development Authority Procedure for large-scale production of astaxanthin from haematococcus
JP2000513938A (ja) 1996-07-01 2000-10-24 アクアサーチ インコーポレイテッド ヘマトコッカス種の成長プロセスの制御方法
WO2003027267A1 (en) 2001-09-26 2003-04-03 Parry Nutraceuticals Ltd. Process to produce astaxanthin from haematococcus biomass
WO2005085465A1 (ja) 2004-03-04 2005-09-15 Suntory Limited アスタキサンチン含有脂質の製造方法
US20150252391A1 (en) 2012-07-27 2015-09-10 East China University Of Science And Technology Method using microalgae for high-efficiency production of astaxanthin

Family Cites Families (16)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
US4683202A (en) 1985-03-28 1987-07-28 Cetus Corporation Process for amplifying nucleic acid sequences
US6680426B2 (en) 1991-01-07 2004-01-20 Auburn University Genetic engineering of plant chloroplasts
WO2002010398A2 (en) 2000-07-31 2002-02-07 Hahn Frederick M Manipulation of genes of the mevalonate and isoprenoid pathways to create novel traits in transgenic organisms
US7329789B1 (en) 2001-10-25 2008-02-12 Phasex Corporation Method for extraction and concentration of carotenoids using supercritical fluids
EP1361280A1 (en) 2002-05-08 2003-11-12 Wageningen University Process for continuous production and extraction of carotenoids from natural sources
WO2006107736A1 (en) 2005-04-01 2006-10-12 Algaen Corporation Extractability and bioavailability of the natural antioxidant astaxanthin from a green alga, haematococcus pluvialis
EP2147113A1 (en) 2007-05-23 2010-01-27 Cognis IP Management GmbH Efficient astaxanthin production strains derived from haematococcus pluvialis
EP1995325A1 (en) 2007-05-23 2008-11-26 Cognis IP Management GmbH Efficient astaxanthin production strains derived from Haematococcus pluvialis
AU2009231810A1 (en) 2008-03-31 2009-10-08 Kuehnle Agrosystems, Inc. Nuclear based expression of genes for production of biofuels and process co-products in algae
US8278090B1 (en) 2008-07-03 2012-10-02 Solazyme, Inc. Heterotrophic cultivation of hydrocarbon-producing microalgae
CH701253A1 (de) 2009-06-12 2010-12-15 Mibelle Ag Verwendung eines Extraktes aus Schneealgen in kosmetischen oder dermatologischen Formulierungen.
ES2732493T3 (es) 2009-09-18 2019-11-22 Phycoil Biotechnology Int Inc Fermentación de microalgas usando iluminación controlada
US9428779B2 (en) * 2010-02-03 2016-08-30 Sapphire Energy, Inc. Transformation of algae for increasing lipid production
USPP23858P3 (en) 2011-04-18 2013-08-27 Karl Y. Biel Chlamydomonas reinhardtii alga variety named ‘DG8-108’
WO2014074772A1 (en) 2012-11-09 2014-05-15 Heliae Development, Llc Mixotrophic, phototrophic, and heterotrophic combination methods and systems
RU2541455C1 (ru) * 2014-10-03 2015-02-10 Институт биологии южных морей им. А.О. Ковалевского Способ культивирования одноклеточной зеленой водоросли haematococcus pluvialis для получения астаксантина

Patent Citations (5)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
US6022701A (en) 1996-02-01 2000-02-08 Ben-Gurion University Of The Negev Research And Development Authority Procedure for large-scale production of astaxanthin from haematococcus
JP2000513938A (ja) 1996-07-01 2000-10-24 アクアサーチ インコーポレイテッド ヘマトコッカス種の成長プロセスの制御方法
WO2003027267A1 (en) 2001-09-26 2003-04-03 Parry Nutraceuticals Ltd. Process to produce astaxanthin from haematococcus biomass
WO2005085465A1 (ja) 2004-03-04 2005-09-15 Suntory Limited アスタキサンチン含有脂質の製造方法
US20150252391A1 (en) 2012-07-27 2015-09-10 East China University Of Science And Technology Method using microalgae for high-efficiency production of astaxanthin

Non-Patent Citations (1)

* Cited by examiner, † Cited by third party
Title
Frontiers in Plant Science,2016,Vol.7,Article 531

Also Published As

Publication number Publication date
EP3478850A1 (en) 2019-05-08
JP2019527052A (ja) 2019-09-26
CA3028227A1 (en) 2018-01-04
JP2022177001A (ja) 2022-11-30
WO2018006068A1 (en) 2018-01-04
US20210198680A1 (en) 2021-07-01
US11034968B2 (en) 2021-06-15
JP2025041773A (ja) 2025-03-26
US20180002711A1 (en) 2018-01-04

Similar Documents

Publication Publication Date Title
JP7549444B2 (ja) 微生物バイオマスのための改善された従属栄養生産方法およびバイオ製品
Ambati et al. Industrial potential of carotenoid pigments from microalgae: Current trends and future prospects
Cezare-Gomes et al. Potential of microalgae carotenoids for industrial application
Christaki et al. Innovative microalgae pigments as functional ingredients in nutrition
Dufossé Pigments, microbial
Johnson et al. Astaxanthin from microbial sources
Jin et al. Microalgal biotechnology: carotenoid production by the green algae Dunaliella salina
Bhosale et al. Microbial xanthophylls
Cysewski et al. Industrial Production of Microalgal Cell‐Mass and Secondary Products‐Species of High Potential: Haematococcus
Waditee-Sirisattha et al. Halophilic microorganism resources and their applications in industrial and environmental biotechnology
Liaqat et al. Mixotrophic cultivation of microalgae for carotenoid production
Chandi et al. Production and characterization of microbial carotenoids as an alternative to synthetic colors: a review
Yusoff et al. Use of microalgae pigments in aquaculture
Schweiggert et al. Carotenoid production by bacteria, microalgae, and fungi
KR101847551B1 (ko) 카로티노이드 계열의 항산화 색소 및 dha를 포함한 바이오오일을 고생산하는 돌연변이 미세조류인 스키조키트리움 속 shg104 균주 및 이의 용도
de Boer Biotechnological production of colorants
Mostafa et al. Microalgae as a source of carotenoids in foods, obstacles and solutions
JP2003523765A (ja) 真核微生物を利用したカロテノイド、キサントフィルおよびアポ−カロテノイドの生産法
Niizawa et al. Astaxanthin production by autotrophic cultivation of Haematococcus pluvialis: A success story
US20130309719A1 (en) Heterotrophic microbial production of xanthophyll pigments
Dufossé Microbial and microalgal carotenoids as colourants and supplements
Doria et al. Influence of light stress on the accumulation of xanthophylls and lipids in Haematococcus pluvialis CCALA 1081 grown under autotrophic or mixotrophic conditions
Xie et al. Astaxanthin biosynthesis for functional food development and space missions
Dufossé Current carotenoid production using microorganisms
Abidin et al. Microalgae as a vaccine delivery system to aquatic organisms

Legal Events

Date Code Title Description
A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20200629

A621 Written request for application examination

Free format text: JAPANESE INTERMEDIATE CODE: A621

Effective date: 20200629

A131 Notification of reasons for refusal

Free format text: JAPANESE INTERMEDIATE CODE: A131

Effective date: 20210713

A601 Written request for extension of time

Free format text: JAPANESE INTERMEDIATE CODE: A601

Effective date: 20211004

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20220111

A02 Decision of refusal

Free format text: JAPANESE INTERMEDIATE CODE: A02

Effective date: 20220426

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20220824

C60 Trial request (containing other claim documents, opposition documents)

Free format text: JAPANESE INTERMEDIATE CODE: C60

Effective date: 20220824

C11 Written invitation by the commissioner to file amendments

Free format text: JAPANESE INTERMEDIATE CODE: C11

Effective date: 20220906

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20220928

A911 Transfer to examiner for re-examination before appeal (zenchi)

Free format text: JAPANESE INTERMEDIATE CODE: A911

Effective date: 20221017

C21 Notice of transfer of a case for reconsideration by examiners before appeal proceedings

Free format text: JAPANESE INTERMEDIATE CODE: C21

Effective date: 20221018

A912 Re-examination (zenchi) completed and case transferred to appeal board

Free format text: JAPANESE INTERMEDIATE CODE: A912

Effective date: 20221216

C211 Notice of termination of reconsideration by examiners before appeal proceedings

Free format text: JAPANESE INTERMEDIATE CODE: C211

Effective date: 20221220

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20240312

A521 Request for written amendment filed

Free format text: JAPANESE INTERMEDIATE CODE: A523

Effective date: 20240628

A61 First payment of annual fees (during grant procedure)

Free format text: JAPANESE INTERMEDIATE CODE: A61

Effective date: 20240830

R150 Certificate of patent or registration of utility model

Ref document number: 7549444

Country of ref document: JP

Free format text: JAPANESE INTERMEDIATE CODE: R150