USPP11065P - Kiwi plant named `Tomua` - Google Patents

Kiwi plant named `Tomua` Download PDF

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Publication number
USPP11065P
USPP11065P US08/996,802 US99680297V US11065P US PP11065 P USPP11065 P US PP11065P US 99680297 V US99680297 V US 99680297V US 11065 P US11065 P US 11065P
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Prior art keywords
fruit
hayward
tomua
cultivar
leaf
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US08/996,802
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Russell G Lowe
Hinga D Marsh
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Horticulture and Food Research Institute of New Zealand Ltd
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    • AHUMAN NECESSITIES
    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01HNEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
    • A01H5/00Angiosperms, i.e. flowering plants, characterised by their plant parts; Angiosperms characterised otherwise than by their botanic taxonomy
    • A01H5/08Fruits
    • AHUMAN NECESSITIES
    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01HNEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
    • A01H6/00Angiosperms, i.e. flowering plants, characterised by their botanic taxonomy
    • YGENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y10TECHNICAL SUBJECTS COVERED BY FORMER USPC
    • Y10TTECHNICAL SUBJECTS COVERED BY FORMER US CLASSIFICATION
    • Y10T292/00Closure fasteners
    • Y10T292/08Bolts
    • Y10T292/0894Spring arm

Definitions

  • Kiwi plants in cultivation are deciduous vines of A. deliciosa. There are more than 60 species in the genus Actinidia originating in China and parts of Asia.
  • the kiwi cultivar ⁇ Hayward ⁇ , developed in New Zealand, is the most widely grown cultivar because of its distinctive green flesh, good flavor and long storage life. Plants are dioecious, so male pollinizers are required as well as female plants to ensure fruit production.
  • Kiwi plants grow vigorously in spring, and rapidly develop a canopy of canes up to 6 m long if not managed correctly. Plants require a mild, warm-temperate climate, free from late spring and early autumn frosts, and need well-drained soils to produce consistent, heavy crops of fruit. Regular irrigation is necessary in dry spells. Flowering occurs in late spring (mid-late November in New Zealand). The distinctive, green-fleshed fruit develop rapidly after pollination and reach 90% of their harvest weight after 3 months. Fruit from the cultivar ⁇ Tomua ⁇ is harvested in early April in New Zealand, while ⁇ Hayward ⁇ is normally harvested during May.
  • the present invention relates to a new and distinctive kiwi plant having an ovoid shaped fruit. More particularly, the new cultivar is designated ⁇ Tomua ⁇ , and is derived from a controlled pollination of A. deliciosa DA02 -- 03, an unpatented male selection of unknown parentage, and ⁇ Hayward ⁇ , also unpatented.
  • the male parent which originated from seeds introduced from China in 1975, was selected as the pollen parent for the crosses because of its very early-flowering characteristics.
  • the seed parent ⁇ Hayward ⁇ was selected for its large size, good flavor and good storage life.
  • the new cultivar of kiwi was created in the course of a plant breeding program which was initiated in 1983 at Hort Research in Te Puke, New Zealand.
  • An early-flowering unpatented A. deliciosa male, DA02 -- 03 was crossed on to the unpatented seed parent ⁇ Hayward ⁇ by Russell Lowe in November 1983. 59 seedlings from this cross were planted out in the field in April 1986.
  • ⁇ Tomua ⁇ was selected by Russell Lowe and Hinga Marsh from 19 female seedlings in this population.
  • the new cultivar can be asexually reproduced as cuttings or by grafting or budding on to seedling or cutting-grown rootstocks of A. deliciosa.
  • ⁇ Tomua ⁇ flowers two weeks ahead of ⁇ Hayward ⁇ , so the usual pollinizers used for ⁇ Hayward ⁇ are ineffective.
  • Two new and unpatented early-flowering pollinizers designated Hortkiwi ⁇ Ranger ⁇ and Hortkiwi ⁇ King ⁇ have been selected as males for use in new plantings of ⁇ Tomua ⁇ .
  • FIG. 1 shows typical fruit on the vine of the cultivar ⁇ Tomua ⁇ .
  • FIG. 2 shows typical fruit on the cultivar ⁇ Tomua ⁇ . Note pointed stylar end.
  • FIG. 3 shows ⁇ Tomua ⁇ fruit in cross-section and profile.
  • FIG. 4 shows ⁇ Hayward ⁇ fruit in cross-section and profile.
  • FIG. 5 shows ⁇ Tomua ⁇ fruit in close up. Note overall similarity to ⁇ Hayward ⁇
  • FIG. 6 shows ⁇ Hayward ⁇ fruit in close up.
  • FIG. 7 shows flowers of the cultivar ⁇ Tomua ⁇ on the vine. Note petals cupped around the ovary.
  • FIG. 8 shows flowers of the cultivar ⁇ Hayward ⁇ on the vine. Note petals in horizontal alignment.
  • FIG. 9 shows mature leaves of the ⁇ Tomua ⁇ vine.
  • FIG. 10 shows mature leaves of the ⁇ Hayward ⁇ vine.
  • Photographs of fruit on the vine were taken just prior to the harvest date. Colors may vary depending upon growing conditions under different climate, soil, and cultivation conditions and the fruit skin color may vary depending upon extent of exposure to direct sunlight.
  • the fruit of ⁇ Tomua ⁇ are ovoid in shape, tapering towards the stylar end, generally circular in cross-section although slightly flattened (FIG. 3).
  • the flesh of ⁇ Tomua ⁇ fruit is medium green like ⁇ Hayward ⁇ with a columella similar to ⁇ Hayward ⁇ (FIG. 4).
  • the hairs on the skin of the fruit of ⁇ Tomua ⁇ are slightly stiffer than those of ⁇ Hayward ⁇ and slightly prickly to the touch, but are very easily removed by rubbing or brushing to reveal a rather shiny skin (FIG. 2).
  • the skin color of ⁇ Tomua ⁇ fruit is a reddish-brown in contrast to that of ⁇ Hayward ⁇ which is a medium brown color.
  • ⁇ Tomua ⁇ fruit are slightly longer than those of ⁇ Hayward ⁇ for an equivalent weight.
  • the length to (maximum) width ratio for Tomua fruit is 7:5 while that of Hayward fruit is 6:5.
  • ⁇ Tomua ⁇ fruit have a higher dry matter content at harvest and are sweeter tasting than ⁇ Hayward ⁇ fruit when ripe.
  • the storage life of ⁇ Tomua ⁇ fruit when held at 0° C. is about 12 weeks while that of ⁇ Hayward ⁇ is up to 25 weeks under ideal conditions.
  • the new cultivar ⁇ Tomua ⁇ is pistillate with imperfect flowers, i.e., only sterile pollen is produced and thus flowers require a pollinizer for fruit production.
  • Two specific pollinizers have been developed for ⁇ Tomua ⁇ and they are named Hortkiwi ⁇ Ranger ⁇ and Hortkiwi ⁇ King ⁇ .
  • Characteristics of the new cultivar by which it differs from the common ⁇ Hayward ⁇ cultivar include earlier flowering and harvest times, longer fruit shape, more easily removed hairs on the skin, more pointed shape at the stylar end (FIG. 2) and reddish-brown colored skin. Horticultural terminology is used in accordance with revised UPOV guidelines for kiwi.
  • ⁇ Tomua ⁇ vines can be grown on the same rootstocks as can ⁇ Hayward ⁇ vines. Rootstocks currently used in New Zealand include A. deliciosa seedlings, ⁇ Hayward ⁇ rooted cuttings and Hortkiwi ⁇ Kaimai ⁇ . ⁇ Tomua ⁇ can also be grown as self-rooted cutting grown plants.
  • the storage life of ⁇ Tomua ⁇ fruit is about half of that of ⁇ Hayward ⁇ fruit under cool storage temperatures of 0° C., but this is not considered a disadvantage as ⁇ Tomua ⁇ fruit can be sold before the main ⁇ Hayward ⁇ crop is harvested.
  • ⁇ Tomua ⁇ fruit soften more rapidly than ⁇ Hayward ⁇ fruit so the consumer can more readily obtain fruit that is in a "ready-to-eat” condition at the start of the kiwi fruit harvest season.
  • ⁇ Tomua ⁇ fruit are sweeter tasting than ⁇ Hayward ⁇ with a tangy balance of sweetness and acidity.

Abstract

A new and distinct kiwi plant of the species Actinidia deliciosa (A.Chev.) C.F. Liang et A.R. Ferguson is described. The cultivar results from a controlled pollination of A. deliciosa DA02-- 03, a male selection of unknown parentage, and the female A. deliciosa `Hayward`, the most widely grown cultivar of kiwi worldwide. Both named parents (DA02-- 03 and `Hayward`) are unpatented cultivars. The new cultivar is distinguished by its early harvest time (about 4 weeks ahead of `Hayward` in NZ), fruit appearance similar to `Hayward`, and moderate plant vigor.

Description

BACKGROUND TO THE INVENTION
Kiwi plants in cultivation are deciduous vines of A. deliciosa. There are more than 60 species in the genus Actinidia originating in China and parts of Asia. The kiwi cultivar `Hayward`, developed in New Zealand, is the most widely grown cultivar because of its distinctive green flesh, good flavor and long storage life. Plants are dioecious, so male pollinizers are required as well as female plants to ensure fruit production.
Kiwi plants grow vigorously in spring, and rapidly develop a canopy of canes up to 6 m long if not managed correctly. Plants require a mild, warm-temperate climate, free from late spring and early autumn frosts, and need well-drained soils to produce consistent, heavy crops of fruit. Regular irrigation is necessary in dry spells. Flowering occurs in late spring (mid-late November in New Zealand). The distinctive, green-fleshed fruit develop rapidly after pollination and reach 90% of their harvest weight after 3 months. Fruit from the cultivar `Tomua` is harvested in early April in New Zealand, while `Hayward` is normally harvested during May.
SUMMARY OF THE INVENTION
The present invention relates to a new and distinctive kiwi plant having an ovoid shaped fruit. More particularly, the new cultivar is designated `Tomua`, and is derived from a controlled pollination of A. deliciosa DA02-- 03, an unpatented male selection of unknown parentage, and `Hayward`, also unpatented. The male parent, which originated from seeds introduced from China in 1975, was selected as the pollen parent for the crosses because of its very early-flowering characteristics. The seed parent `Hayward` was selected for its large size, good flavor and good storage life.
The new cultivar of kiwi was created in the course of a plant breeding program which was initiated in 1983 at Hort Research in Te Puke, New Zealand. An early-flowering unpatented A. deliciosa male, DA02-- 03, was crossed on to the unpatented seed parent `Hayward` by Russell Lowe in November 1983. 59 seedlings from this cross were planted out in the field in April 1986. By December 1990 most plants had flowered and `Tomua` was selected by Russell Lowe and Hinga Marsh from 19 female seedlings in this population. `Tomua` fruit matured 4 weeks ahead of `Hayward` and had acceptable fruit size, a good flavor and appeared to have commercial potential.
The new cultivar can be asexually reproduced as cuttings or by grafting or budding on to seedling or cutting-grown rootstocks of A. deliciosa. Trial plantings, established in 1989 at Te Puke, with plants grafted on to clonal `Hayward` rootstocks, have shown that the unique combination of characteristics come true to form and are established and transmitted through succeeding asexual propagations. `Tomua` flowers two weeks ahead of `Hayward`, so the usual pollinizers used for `Hayward` are ineffective. Two new and unpatented early-flowering pollinizers designated Hortkiwi `Ranger` and Hortkiwi `King` have been selected as males for use in new plantings of `Tomua`.
BRIEF DESCRIPTION OF THE ILLUSTRATIONS
FIG. 1 shows typical fruit on the vine of the cultivar `Tomua`.
FIG. 2 shows typical fruit on the cultivar `Tomua`. Note pointed stylar end.
FIG. 3 shows `Tomua` fruit in cross-section and profile.
FIG. 4 shows `Hayward` fruit in cross-section and profile.
FIG. 5 shows `Tomua` fruit in close up. Note overall similarity to `Hayward`
FIG. 6 shows `Hayward` fruit in close up.
FIG. 7 shows flowers of the cultivar `Tomua` on the vine. Note petals cupped around the ovary.
FIG. 8 shows flowers of the cultivar `Hayward` on the vine. Note petals in horizontal alignment.
FIG. 9 shows mature leaves of the `Tomua` vine.
FIG. 10 shows mature leaves of the `Hayward` vine.
Photographs of fruit on the vine were taken just prior to the harvest date. Colors may vary depending upon growing conditions under different climate, soil, and cultivation conditions and the fruit skin color may vary depending upon extent of exposure to direct sunlight.
The fruit of `Tomua` are ovoid in shape, tapering towards the stylar end, generally circular in cross-section although slightly flattened (FIG. 3). The flesh of `Tomua` fruit is medium green like `Hayward` with a columella similar to `Hayward` (FIG. 4). The hairs on the skin of the fruit of `Tomua` are slightly stiffer than those of `Hayward` and slightly prickly to the touch, but are very easily removed by rubbing or brushing to reveal a rather shiny skin (FIG. 2). The skin color of `Tomua` fruit is a reddish-brown in contrast to that of `Hayward` which is a medium brown color. `Tomua` fruit are slightly longer than those of `Hayward` for an equivalent weight. The length to (maximum) width ratio for Tomua fruit is 7:5 while that of Hayward fruit is 6:5. `Tomua` fruit have a higher dry matter content at harvest and are sweeter tasting than `Hayward` fruit when ripe. The storage life of `Tomua` fruit when held at 0° C. is about 12 weeks while that of `Hayward` is up to 25 weeks under ideal conditions.
BOTANICAL DESCRIPTION OF THE PLANT
The new cultivar `Tomua` is pistillate with imperfect flowers, i.e., only sterile pollen is produced and thus flowers require a pollinizer for fruit production. Two specific pollinizers have been developed for `Tomua` and they are named Hortkiwi `Ranger` and Hortkiwi `King`. Characteristics of the new cultivar by which it differs from the common `Hayward` cultivar include earlier flowering and harvest times, longer fruit shape, more easily removed hairs on the skin, more pointed shape at the stylar end (FIG. 2) and reddish-brown colored skin. Horticultural terminology is used in accordance with revised UPOV guidelines for kiwi.
______________________________________
TABLE OF CHARACTERISITCS
Characters of the `Hayward` comparison cultivar are noted in  bracket!
opposite that character only when significantly different.
All dimensions in millimeters, weights in grams.
               Tomua      Hayward
______________________________________
PLANT
Plant: sex expression
                 female
Plant: ploidy    hexaploid (2n =
                 6x = 174)
Plant: vigor     medium
Young shoot: hairs
                 present
Young shoot: density of
                 medium
hairs
Young shoot: type of hairs
                 velutinous
Young shoot: anthocyanin
                 medium
coloration of growing tip
young shoot: anthocyanin
                 weak
coloration of leaf axil
STEM
Stem: diameter   medium - Mean
                 10.8 mm (Range
                 10.2-12.3)
Stem: dormant bud diameter
                 2.55 mm
                 (1.7-3.1)
Stem: color on upper side
                 red-brown
of shoot
Stem: hairs      present
Stem: conspicuousness of
                 conspicuous
lenticels
Stem: number of lenticels
                 medium
Stem: color of lenticels
                 brownish-
                 white
Stem: size of bud support
                 medium
Stem: visibility of bud
                 inconspicuous
(dormant canes)
Stem: number of hairs
                 few
visible on bud
(dormant canes)
Stem: leaf scar  medium
LEAF
Leaf: general shape of
                 round         very
blade                         broadly
                              ovate!
Leaf: length     140 mm
                 (120-160 mm)
Leaf: width      149 mm
                 (100-170 mm)
Leaf: shape of tip of blade
                 mucronate
Leaf: shape of base of blade
                 cordate
Leaf: arrangement of leaf
                 overlapping
bases
Leaf: puckering/blistering
                 medium
on upper side of blade
Leaf: margin     ciliate
Leaf: green color of upper
                 medium
side of blade
Leaf: glossiness of upper
                 medium
surface of blade
Leaf: color of lower side
                 light
of blade         green
Leaf: glaucousness (lower
                 absent
side of blade)
Leaf: petiole length
                 87 mm
                 (52-150 mm)
Leaf: hears on petiole
                 present
Leaf: density of hairs on
                 medium
petiole
Leaf: anthocyanin coloration
                 weak          medium!
on upper side of petiole
FLOWER (Measurements taken
from a sample of 20 flowers)
Inflorescence: predom-
                 one
inant number of flowers
Pedicel: length  medium -
                 36.9 mm
                 (31.0-43.0 mm)
Pedicel: hairs   present
Pedicel: length of hairs
                 medium
Flower: number of sepals
                 >5
Flower: color of sepals
                 brown
Flower: diameter (terminal
                 large -
or king flower when fully
                 63.3 mm
open)            (47.8-70.6 mm)
Flower: petal length
                 30.0 mm
                 (25.3-34.3
                 mm)
Flower: petal width
                 22.7 mm
                 (18.5-29.6
                 mm)
Flower: petal length/
                 1.33 mm
width ratio      (1.11-1.48
                 mm)
Flower: mean number of
                 7 (6-8)
petals per flower
Flower: number of flowers
                 15 out of
with more than six petals
                 20 flowers
Flower: arrangement of
                 overlapping
petals
Flower: petal shoulder
                 present
Flower: primary color of
                 white
petals on upper side (when
fully open)
Flower: type of coloration
                 uniform color,
of petals        i.e., not bi-
                 colored
Flower: number of styles
                 37.3 (31-43)
Flower: attitude of styles
                 semi-erect
Flower: amount of hair on
                 strongly
ovary            expressed
FRUIT Measurements are from
10 fruit sampes
Fruit: overall size
                 medium 91 g   large!
Fruit: mean length
                 73.9 mm
                 (69.1-79.1
                 mm)
Fruit: width (max.)
                 51.56 mm
                 (47.4-55.0
                 mm)
Fruit: width (min.)
                 44.9 mm
                 42.8-47.3
                 mm)
Fruit: locule number
                 36.9 (33-40)
Fruit: general shape
                 ovoid (tapers
                 toward stylar end)
Fruit: cross-section at
                 elliptical    oblate!
median
Fruit: general shape of
                 raised        flat!
stylar end
Fruit: shape of shoulder
                 rounded       almost
on salk end                   square!
Fruit: skin color at harvest
                 reddish-      brown!
(fruit still hard)
                 brown
Fruit: skin color change
                 absent
during ripening
Fruit: skin color at
                 reddish-      brown!
maturity for consumption
                 brown
Fruit: hairs     present
Fruit: density of hairs
                 medium
Fruit: type of hair
                 bristly       hirsute!
Fruit: concentration of hairs
                 uniform
Fruit: adherenece of hairs to
                 weak          strong!
skin (when rubbed)
Fruit: core (columella)
                 medium to     large!
diameter (at largest dia.)
                 large
Fruit: core shape (in cross
                 elliptical
section)
Fruit: core woody spike
                 present
Fruit: prominence of core
                 medium
woody spike
Fruit: outer pericarp color
                 light green
at maturity for consumption
Fruit: inner pericarp color
                 green
(locules) at maturity for
consumption
Fruit: core color at maturity
                 greenish
                 white
Fruit: soluble solids
                 medium -
concentration SSC
                 14.3%
(Brix level) at maturity
                 (13.5-15.1%)
for consumption
Fruit: Vitamin C content at
                 medium
harvest          (100-125 mg/
                 100 g fresh
                 weight)
Fruit: seed color at maturity
                 dark
                 brown
Fruit: seed color when dry
                 brown
EVENTS (at Te Puke,
New Zealand)
Time of vegetative
                 early (late
budbreak         August to first
                 week of Sept.)
Time of beginning of
                 medium (second)
                               two weeks
flowering        to third week
                              later!
                 of Sept.)
Time of harvest at 6.2%
                 medium (first
                               four weeks
SSC              weak of April)
                              later!
______________________________________
HORTICULTURAL CHARACTERISTICS
Details below relate to observations made on plants grafted on `Hayward` clonal rootstocks (rooted cuttings) growing at Te Puke Research Centre, New Zealand.
`Tomua` vines can be grown on the same rootstocks as can `Hayward` vines. Rootstocks currently used in New Zealand include A. deliciosa seedlings, `Hayward` rooted cuttings and Hortkiwi `Kaimai`. `Tomua` can also be grown as self-rooted cutting grown plants.
In Te Puke, from 1993-1997 the `Tomua` harvest time has been consistently 4-5 weeks ahead of `Hayward` using the criteria of a Soluble Solids Concentration (SSC) of 6.2% to begin harvest.
The storage life of `Tomua` fruit is about half of that of `Hayward` fruit under cool storage temperatures of 0° C., but this is not considered a disadvantage as `Tomua` fruit can be sold before the main `Hayward` crop is harvested.
`Tomua` fruit soften more rapidly than `Hayward` fruit so the consumer can more readily obtain fruit that is in a "ready-to-eat" condition at the start of the kiwi fruit harvest season. `Tomua` fruit are sweeter tasting than `Hayward` with a tangy balance of sweetness and acidity.
Yield data: `Tomua` vines carry lower yields of Class 1 (export grade) fruit than `Hayward` when plants of equivalent ages are compared, mainly because fruit of `Tomua` are smaller on average than those of `Hayward`.
Three years of data from a replicated trial planting containing `Tomua` and `Hayward`, planted in 1989 at Te Puke is presented below in Table 1.
              TABLE 1
______________________________________
TOMUA AND HAYWARD YIELDS
Fruit number   Trays (3.6 kg)
                            Size (grams)
Year  Tomua   Hayward  Tomua Hayward
                                    Tomua Hayward
______________________________________
1996  938     708      23    22     92    117
1997  909     1331     23    35     93    96
1998  1149    1020     24    30     89    111
Means 999     1020     23    29     91    108
______________________________________
Fruit flesh and skin color measurement.
RHS Colour Chart, the Royal Horticultural Society, London
______________________________________
1966.
       Flesh Color at maturity:
Outer pericarp
              138B-138D   Hayward 138B-138C!
Tomua
Fruit core:
Tomua         155-155B    Hayward 155A!
Seed color (in fruit):
              200A
Seed color (dry seed):
              165A-165B
Fruit skin at maturity:
Tomua         165B        Hayward 164B!
Leaf color: mature leaf
after petal fall
Upper side of leaf
              137A or
Tomua         147A
Lower side of leaf
              147B-147C
Tomua
Petiole color:
exposed side  178B
Flower petals:
main body     155D
base          154C-154D
Plant Stem:
exposed side: 165A
Lenticel color:
               164B-164C.
______________________________________

Claims (1)

We claim:
1. A new and distinct kiwi plant of the species A. deliciosa substantially as described and illustrated in the specification above, characterised by early harvest date compared to `Hayward`, medium green flesh, reddish-brown skin, easily removable hairs, and a slight pointed stylar end.
US08/996,802 1997-12-23 1997-12-23 Kiwi plant named `Tomua` Expired - Lifetime USPP11065P (en)

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Cited By (1)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
USPP22276P3 (en) 2009-06-30 2011-11-29 Zespri Group Limited Kiwifruit plant named ‘ZESH004’

Non-Patent Citations (2)

* Cited by examiner, † Cited by third party
Title
UPOV CD ROM Kiwi plant named Tomua , PBR KIW009, New Zealand, May 1995. *
UPOV CD-ROM Kiwi plant named `Tomua`, PBR KIW009, New Zealand, May 1995.

Cited By (1)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
USPP22276P3 (en) 2009-06-30 2011-11-29 Zespri Group Limited Kiwifruit plant named ‘ZESH004’

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Owner name: HORTICULTURE AND FOOD RESEARCH INSTITUTE OF NEW ZE

Free format text: ASSIGNMENT OF ASSIGNORS INTEREST;ASSIGNORS:LOWE, RUSSELL G.;MARSH, HINGA D.;REEL/FRAME:012547/0921

Effective date: 20010110