CN103688900B - A kind of Japanese instar chickling plumage look that can make is accurately from the producing method for seed of other male and female - Google Patents
A kind of Japanese instar chickling plumage look that can make is accurately from the producing method for seed of other male and female Download PDFInfo
- Publication number
- CN103688900B CN103688900B CN201310310478.XA CN201310310478A CN103688900B CN 103688900 B CN103688900 B CN 103688900B CN 201310310478 A CN201310310478 A CN 201310310478A CN 103688900 B CN103688900 B CN 103688900B
- Authority
- CN
- China
- Prior art keywords
- plumage
- cock
- hen
- iicocoss
- female
- Prior art date
- Legal status (The legal status is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the status listed.)
- Expired - Fee Related
Links
- 238000000034 method Methods 0.000 title claims abstract description 21
- 241000287828 Gallus gallus Species 0.000 claims abstract description 23
- 210000003746 feather Anatomy 0.000 claims abstract description 21
- BQCADISMDOOEFD-UHFFFAOYSA-N Silver Chemical compound [Ag] BQCADISMDOOEFD-UHFFFAOYSA-N 0.000 claims description 22
- 229910052709 silver Inorganic materials 0.000 claims description 22
- 239000004332 silver Substances 0.000 claims description 22
- 238000012360 testing method Methods 0.000 claims description 13
- 108090000623 proteins and genes Proteins 0.000 claims description 11
- 108700028369 Alleles Proteins 0.000 claims description 7
- 239000000835 fiber Substances 0.000 claims description 3
- 238000012797 qualification Methods 0.000 claims description 3
- 210000003765 sex chromosome Anatomy 0.000 claims description 3
- 239000011435 rock Substances 0.000 claims description 2
- 230000013011 mating Effects 0.000 description 3
- FFRBMBIXVSCUFS-UHFFFAOYSA-N 2,4-dinitro-1-naphthol Chemical compound C1=CC=C2C(O)=C([N+]([O-])=O)C=C([N+]([O-])=O)C2=C1 FFRBMBIXVSCUFS-UHFFFAOYSA-N 0.000 description 2
- 108700011325 Modifier Genes Proteins 0.000 description 2
- 238000009395 breeding Methods 0.000 description 2
- 238000012850 discrimination method Methods 0.000 description 2
- PCHJSUWPFVWCPO-UHFFFAOYSA-N gold Chemical compound [Au] PCHJSUWPFVWCPO-UHFFFAOYSA-N 0.000 description 2
- 229910052737 gold Inorganic materials 0.000 description 2
- 239000010931 gold Substances 0.000 description 2
- 238000007689 inspection Methods 0.000 description 2
- 230000008774 maternal effect Effects 0.000 description 2
- 210000001550 testis Anatomy 0.000 description 2
- 230000001488 breeding effect Effects 0.000 description 1
- 230000000694 effects Effects 0.000 description 1
- 238000005516 engineering process Methods 0.000 description 1
- 230000007774 longterm Effects 0.000 description 1
- 235000013372 meat Nutrition 0.000 description 1
- 210000001672 ovary Anatomy 0.000 description 1
- 238000011160 research Methods 0.000 description 1
- 238000005070 sampling Methods 0.000 description 1
- 238000012549 training Methods 0.000 description 1
Landscapes
- Measuring Or Testing Involving Enzymes Or Micro-Organisms (AREA)
Abstract
The present invention relates to a kind of Japanese instar chickling plumage look that can make accurately from the producing method for seed of other male and female, the method comprises the following steps: 1) hybridize to obtain F1 pure white feather cock; 2) F1 white plumage cock backcrosses to yellow plumage this hen public, and select and remain white plumage black tail cock and Bai Yu black tail hen; 3) white plumage black tail male and female chicken from numerous and be measured.The Japanese instar chickling that the method is produced, all can plumage look from other male and female, and all reach 100% from not other rate.
Description
Technical field
The present invention relates to a kind of chicken breeding method, particularly relate to a kind of Japanese instar chickling plumage look that can make accurately from the producing method for seed of other male and female.
Background technology
Since laying hen and meat chicken enter scale and Virtual production, laying hen and kind chicken all require to differentiate male and female when an age in days, to alleviate feeding cost in the future.Current male and female discrimination method has two kinds, and one is anal identification method, and two is utilize plumage speed or plumage look from other.Anal identification method not only requires that discriminating personnel carry out long-term professional training, and labour intensity is large, resolution has varied with each individual, and height has low, and misoperation easily hinders chicken.One Japanese instar chickling male and female is divided from other from not having plumage look and plumage speed, wherein the most simply, be utilize sex-linked golden feather genes (s) silver color feather genes (S) to carry out plumage look from other the most fast, wherein s and S is allelomorph, silver color plumage is dominant relative to golden plumage, but and not exclusively.As shown in Figure 1, its general principle and flow process as follows:
First set up respectively two strains being with isozygoty golden plumage and the silver color plumage that isozygotys, and do with silver color plumage system maternal, golden plumage system makes male parent, and in one age in days offspring, white person is public young bird, and golden person is female young.
Research shows; because silver color plumage is incomplete dominance to golden plumage; so when lacking modifier gene; utilize the male and female of gold and silver look feather genes from rate is not high; usually can only reach between 90-95%; even lower (Kang Xiangtao etc. 1995, peaceful China 2008), cannot reach the effect that scale utilizes.
Applicant has applied for that (application number: 201010183909.7 applyings date: 2010-05-27) discloses a kind of Japanese instar chickling plumage look that can make accurately from the producing method for seed of other male and female; the method comprises the following steps: 1) proof gold look Colombia plumage cock and standard variety White Rock hybridize, and hybridizes to obtain F1 pure white feather cock; 2) F1 pure white feather cock backcrosses to yellow plumage this hen public, and select and remain white plumage black tail cock and Bai Yu black tail hen; 3) silvery white plumage black tail cock and Bai Yu black tail hen are from numerous, and hen stays silvery white plumage black tail person, eliminate golden plumage individual; Cock stays typical Colombia plumage type individual, and carries out being measured with silvery white plumage black tail hen, eliminates the heterozygosis cock occurring non-silvery white typical case Colombia plumage type individuality in offspring.4) silvery white plumage black tail hen and red Yufu system or the mating of yellow Yufu system of being with golden feather genes, the Japanese instar chickling produced, white plumage type is cock, and golden yellow plumage is hen; 100% is reached from not other rate.But there is a difficult problem in this patent, in step 3), cock stays typical Colombia plumage type individual, genotype is iiCoCoSS or iiCoCoSs, and carry out being that iiCoCoSW hen is measured with genotype, eliminate individual iiCoCoSs and iiCocoSS of heterozygosis cock occurring non-silvery white typical case Colombia plumage type individuality in offspring, be difficult to obtain the individual iiCocoSS of heterozygosis cock.
Summary of the invention
In order to solve above-mentioned technical problem, the object of this invention is to provide a kind of Japanese instar chickling plumage look that can make accurately from the producing method for seed of other male and female, the Japanese instar chickling that the method is produced, all can plumage look from other male and female, and all reach 100% from not other rate.
In order to realize above-mentioned technical purpose, present invention employs following technical scheme:
A Japanese instar chickling plumage look can be made accurately from a producing method for seed for other male and female, and the method comprises the following steps:
1) F1 pure white feather cock is hybridized to obtain, as shown in Figure 2;
2) F1 white plumage cock backcrosses to yellow plumage this hen public, and select and remain white plumage black tail cock and Bai Yu black tail hen, as shown in Figure 3;
3) white plumage black tail male and female chicken from numerous and be measured: male and female chicken leaves white plumage black tail person for the first time after numerous, obtains iiCo-S-cock and iiCo-SW hen; Utilize fried dough twist chicken, its genotype is public iicocoss, female iicocosW; All individualities are measured, obtain single heterozygosis cock iiCoCoSs and carry out secondary from numerous with the colony of the hen iiCoCoSW that isozygotys, equally only stay white plumage black tail individual, then to the homozygosity in mature rooster qualification S site, and finally set up the target group of isozygotying completely: cock is iiCoCoSS, and hen is iiCoCoSW; Concrete steps are as follows:
First test cross of joining fried dough twist hen iicocosW from numerous cock, in order to the mono-heterozygosis cock of the iiCoCoSs that selects and remain; Superseded SS isozygotys cock: offspring chick is all silver color plumage; Stay coloured young bird of Ss heterozygosis parent cock to observe further: if there is not obvious numb plumage proterties in coloured young bird, chick has the black flower of sheet, and Adult Chicken is fried dough twist look, its parent cock is the mono-heterozygous individual of iiCoCoSs; If the numb plumage proterties of about 50% individual appearance in coloured young bird, then parent cock is the two heterozygous individual of iiCocoSs; Secondary is from numerous cock with isozygotying golden plumage hen test cross and seek obtain iiCoCoSS homozygous individual group, and offspring is all the parent cock of silver color plumage is iiCoCoSS, otherwise is iiCoCoSs; Silver color plumage hen is without the need to test cross again;
The test cross of fiber crops plumage cock iicocoss dialogue plumage black tail hen iiCo-SW: public young bird is silver color, and female young bird is coloured plumage; If had an appointment in female young bird, the individuality of 50% shows numb plumage proterties, chick has the black flower of sheet, and Adult Chicken is fried dough twist look, and parent hen is the iiCocoSW of heterozygosis, individual without obvious numb plumage proterties in female young bird, without obvious black plumage or only have the parent of banded black decorative pattern to be then iiCoCoSW;
As shown in Figure 4;
Above-mentioned I and i is allelomorph, and I is dominant white feather genes, and i is not then with dominant white feather genes; With the individuality of Co, under the background not being pure white feather, be usually expressed as Colombia's plumage type; S is silver color plumage, and s is its golden plumage allelomorph; W-Z is sex chromosome, and wherein cock is ZZ, hen ZW.
As preferably, isozygoty golden plumage hen H33, fried dough twist hen or island, standard variety Lip river of described step 3) is red.
The silver color plumage of the band Colombia modifier gene that the present invention utilizes the art of this patent to set up is maternal, can with red Yufu system (as red treasured), yellow Yufu system (plumage landrace as yellow in the major part) mating of being with golden feather genes, the Japanese instar chickling produced, all can plumage look from other male and female, and all reach 100% from not other rate.
Accompanying drawing explanation
Fig. 1 is the flow chart of existing male and female discrimination method.
Fig. 2 is that the flow chart of F1 pure white feather cock is hybridized to obtain in the present invention.
Fig. 3 is the flow chart that F1 of the present invention white plumage cock backcrosses to yellow plumage this hen public.
Fig. 4 is the flow chart that the present invention finally sets up the target group of isozygotying completely.
Embodiment
A Japanese instar chickling plumage look can be made accurately from a producing method for seed for other male and female, and the method comprises the following steps:
1) as shown in Figure 2, F1 pure white feather cock is hybridized to obtain
2) as shown in Figure 3, F1 white plumage cock backcrosses to yellow plumage this hen public, and select and remain white plumage black tail cock and Bai Yu black tail hen (accounting for 1/4 of offspring's sum).
3) white plumage black tail male and female chicken from numerous and be measured: male and female chicken leaves white plumage black tail person for the first time after numerous, obtains iiCo-S-cock and iiCo-SW hen.(its genotype is public iicocoss to utilize fried dough twist chicken, female iicocosW) all individualities are measured, the colony obtaining single heterozygosis cock (iiCoCoSs) and hen of isozygotying (iiCoCoSW) carries out secondary from numerous, equally only stay white plumage black tail individual, then to the homozygosity in mature rooster qualification S site, and finally set up target group's (cock is iiCoCoSS, and hen is iiCoCoSW) of isozygotying completely.As shown in Figure 4, concrete steps are as follows:
First test cross of joining fried dough twist hen iicocosW from numerous cock, in order to the mono-heterozygosis cock of the iiCoCoSs that selects and remain.Superseded SS isozygotys cock: offspring chick is all silver color plumage; Ss heterozygosis parent cock is stayed (to have an appointment 50% in offspring for coloured young bird, all the other 50% are white young) coloured young bird observe further: if do not occur that in coloured young bird (chick has the black flower of sheet to obvious numb plumage proterties, Adult Chicken is fried dough twist look), its parent cock is the mono-heterozygous individual of iiCoCoSs; If the numb plumage proterties of about 50% individual appearance in coloured young bird, then parent cock is the two heterozygous individual of iiCocoSs.Secondary can with any golden plumage hen of isozygotying (as red with H33, fried dough twist hen or island, standard variety Lip river from numerous cock, we are with H33) test cross and seek obtain iiCoCoSS homozygous individual group (offspring is all the parent cock of silver color plumage is iiCoCoSS, otherwise is iiCoCoSs).Silver color plumage hen is without the need to test cross again.
The test cross of fiber crops plumage cock iicocoss dialogue plumage black tail hen iiCo-SW: public young bird is silver color, and female young bird is coloured plumage.If had an appointment in female young bird, the individuality of 50% shows numb plumage proterties (chick has the black flower of sheet, Adult Chicken is fried dough twist look), parent hen is the iiCocoSW of heterozygosis, is then iiCoCoSW without the parent of obvious numb plumage proterties individuality (without obvious black plumage or only have banded black decorative pattern) in female young bird.
Note: I and i is allelomorph, and I is dominant white feather genes, and i is not then with dominant white feather genes; With the individuality of Co, under the background not being pure white feather, be usually expressed as Colombia's plumage type; S is silver color plumage, and s is its golden plumage allelomorph; W-Z is sex chromosome, and wherein cock is ZZ, hen ZW.
The hen utilizing above-mentioned producing method for seed to obtain respectively with red plumage cock, the mating of yellow plumage cock, hatch results is as following table
To golden yellow type breeding chickling to growing up, be hen without exception; The offspring chick of white plumage type is not raised, but directly carries out sampling and cut open inspection: from the offspring joining red plumage cock and join yellow plumage cock, respectively get 200 immediately, check whether to there is testis or ovary after cuing open inspection.As a result, regard as all 400 chick of white plumage type, all have testis, thus confirm that 100% is cock.In test, male and female is from other rate of accuracy reached 100%.
Claims (2)
1. a Japanese instar chickling plumage look can be made accurately from a producing method for seed for other male and female, it is characterized in that the method comprises the following steps:
1) F1 pure white feather cock is hybridized to obtain: by H33: cock genotype is iiCoCoss and standard variety White Rock: hen genotype is that IIcocoSW is hybridized and obtained F1 cock;
2) F1 white plumage cock genotype is IiCocoSs is that iiCoCosW backcrosses to H33 yellow plumage hen genotype, and 1/4 white plumage black tail cock genotype of selecting and remain is iiCo-Ss and 1/4 plumage black tail hen iiCo-SW in vain:
3) white plumage black tail male and female chicken from numerous and top-cross: male and female chicken leaves white plumage black tail person for the first time after numerous, obtains iiCo-S-cock and iiCo-SW hen; Utilize fried dough twist chicken, its genotype is public iicocoss, female iicocosW; Top-cross is carried out to all individualities, obtain single heterozygosis cock iiCoCoSs and carry out secondary from numerous with the colony of the hen iiCoCoSW that isozygotys, equally only stay white plumage black tail individual, then to the homozygosity in mature rooster qualification S site, and finally set up the target group of isozygotying completely: cock is iiCoCoSS, and hen is iiCoCoSW; Concrete steps are as follows:
First test cross of joining fried dough twist hen iicocosW from numerous cock, in order to the mono-heterozygosis cock of the iiCoCoSs that selects and remain; Superseded SS isozygotys cock: offspring chick is all silver color plumage; Stay coloured young bird of Ss heterozygosis parent cock to observe further: if there is not obvious numb plumage proterties in coloured young bird, chick has the black flower of sheet, and Adult Chicken is fried dough twist look, its parent cock is the mono-heterozygous individual of iiCoCoSs; If the numb plumage proterties of about 50% individual appearance in coloured young bird, then parent cock is the two heterozygous individual of iiCocoSs; Secondary is from numerous cock with isozygotying golden plumage hen test cross and seek obtain iiCoCoSS homozygous individual group, and offspring is all the parent cock of silver color plumage is iiCoCoSS, otherwise is iiCoCoSs; Silver color plumage hen is without the need to test cross again;
The test cross of fiber crops plumage cock iicocoss dialogue plumage black tail hen iiCo-SW: public young bird is silver color, and female young bird is coloured plumage; If have the individuality of 50% to show numb plumage proterties in female young bird, chick has the black flower of sheet, and Adult Chicken is fried dough twist look, and parent hen is the iiCocoSW of heterozygosis, individual without obvious numb plumage proterties in female young bird, without obvious black plumage or only have the parent of banded black decorative pattern to be then iiCoCoSW;
Above-mentioned I and i is allelomorph, and I is dominant white feather genes, and i is not then with dominant white feather genes; With the individuality of Co, under the background not being pure white feather, be usually expressed as Colombia's plumage type; S is silver color plumage, and s is its golden plumage allelomorph; W-Z is sex chromosome, and wherein cock is ZZ, hen ZW.
2. a kind of Japanese instar chickling plumage look according to claim 1 that can make is accurately from the producing method for seed of other male and female, it is characterized in that isozygoty golden plumage hen H33, fried dough twist hen or island, standard variety Lip river of step 3) is red.
Priority Applications (1)
| Application Number | Priority Date | Filing Date | Title |
|---|---|---|---|
| CN201310310478.XA CN103688900B (en) | 2013-07-23 | 2013-07-23 | A kind of Japanese instar chickling plumage look that can make is accurately from the producing method for seed of other male and female |
Applications Claiming Priority (1)
| Application Number | Priority Date | Filing Date | Title |
|---|---|---|---|
| CN201310310478.XA CN103688900B (en) | 2013-07-23 | 2013-07-23 | A kind of Japanese instar chickling plumage look that can make is accurately from the producing method for seed of other male and female |
Publications (2)
| Publication Number | Publication Date |
|---|---|
| CN103688900A CN103688900A (en) | 2014-04-02 |
| CN103688900B true CN103688900B (en) | 2016-01-06 |
Family
ID=50350693
Family Applications (1)
| Application Number | Title | Priority Date | Filing Date |
|---|---|---|---|
| CN201310310478.XA Expired - Fee Related CN103688900B (en) | 2013-07-23 | 2013-07-23 | A kind of Japanese instar chickling plumage look that can make is accurately from the producing method for seed of other male and female |
Country Status (1)
| Country | Link |
|---|---|
| CN (1) | CN103688900B (en) |
Families Citing this family (4)
| Publication number | Priority date | Publication date | Assignee | Title |
|---|---|---|---|---|
| CN104957085A (en) * | 2015-06-12 | 2015-10-07 | 广西南丹益康牧业开发有限公司 | Broiler breeding stealing preventing method |
| CN106070046A (en) * | 2016-06-29 | 2016-11-09 | 来宾市忻城县润华牧业有限责任公司 | A kind of by nascent chickling plumage type discriminating black plumage black skin type Gallus Domesticus property method for distinguishing |
| CN111345267B (en) * | 2020-04-08 | 2021-11-19 | 山东省农业科学院家禽研究所（山东省无特定病原鸡研究中心） | Breeding method of hybrid line of special laying hens with feather color double-self-distinguishing male and female black-feather pink shells and high yield |
| CN111838071B (en) * | 2020-08-07 | 2022-05-03 | 江苏省家禽科学研究所 | Method for breeding pigeons with black tail and/or phoenix head characters |
Citations (5)
| Publication number | Priority date | Publication date | Assignee | Title |
|---|---|---|---|---|
| CN2579170Y (en) * | 2002-11-28 | 2003-10-15 | 浙江大学 | Automatic identifier for sex of chick |
| CN1623380A (en) * | 2003-12-03 | 2005-06-08 | 康相涛 | Method of raising black-bone chicken |
| CN1669411A (en) * | 2005-04-13 | 2005-09-21 | 中国农业科学院家禽研究所 | Indigenous breed layers mark matching technology |
| CN1843104A (en) * | 2006-01-23 | 2006-10-11 | 尹华贵 | Male and female identification method from chicken shank color or sex-linked feather |
| CN101874479A (en) * | 2010-05-27 | 2010-11-03 | 楼梦良 | Breeding method capable of realizing feather color-based accurate self identification of sex of one-day chicken |
Family Cites Families (3)
| Publication number | Priority date | Publication date | Assignee | Title |
|---|---|---|---|---|
| RO117750B1 (en) * | 2002-04-15 | 2002-07-30 | Avicola Bucureşti S.A. | Line of hens (gallus domesticus) of mixed 5a red rhode- island breeds |
| JP2011142866A (en) * | 2010-01-15 | 2011-07-28 | Ryosuke Taniguchi | Method for identifying sex of fertilized egg based on outline thereof |
| KR20130023909A (en) * | 2011-08-30 | 2013-03-08 | 경남과학기술대학교 산학협력단 | Dna probe for gender determination of chick and method for gender determination of chick by the probe using feather |
-
2013
- 2013-07-23 CN CN201310310478.XA patent/CN103688900B/en not_active Expired - Fee Related
Patent Citations (5)
| Publication number | Priority date | Publication date | Assignee | Title |
|---|---|---|---|---|
| CN2579170Y (en) * | 2002-11-28 | 2003-10-15 | 浙江大学 | Automatic identifier for sex of chick |
| CN1623380A (en) * | 2003-12-03 | 2005-06-08 | 康相涛 | Method of raising black-bone chicken |
| CN1669411A (en) * | 2005-04-13 | 2005-09-21 | 中国农业科学院家禽研究所 | Indigenous breed layers mark matching technology |
| CN1843104A (en) * | 2006-01-23 | 2006-10-11 | 尹华贵 | Male and female identification method from chicken shank color or sex-linked feather |
| CN101874479A (en) * | 2010-05-27 | 2010-11-03 | 楼梦良 | Breeding method capable of realizing feather color-based accurate self identification of sex of one-day chicken |
Non-Patent Citations (2)
| Title |
|---|
| 肉用仔鸡自别雌雄配套系的培育;杜炳旺等;《家禽》;19860501(第2期);第34-35页 * |
| 鸡性别鉴定技术的研究概况;林金杏;《中国家禽》;20080405;第30卷(第7期);第35-38页 * |
Also Published As
| Publication number | Publication date |
|---|---|
| CN103688900A (en) | 2014-04-02 |
Similar Documents
| Publication | Publication Date | Title |
|---|---|---|
| CN101874479A (en) | Breeding method capable of realizing feather color-based accurate self identification of sex of one-day chicken | |
| CN103688900B (en) | A kind of Japanese instar chickling plumage look that can make is accurately from the producing method for seed of other male and female | |
| CN108192981B (en) | The white red plumage reason mutated-genotype identification of Leghorn and red feather pink-egg layer chickens breed system breeding method | |
| CN100456926C (en) | Male and female identification method from chicken shank color or sex-linked feather | |
| CN104285880A (en) | Hybrid breed production mode of black feather chicken commercial line | |
| CN104357553A (en) | Pelteobagrus fulvidraco microsatellite family identification method | |
| CN101904318A (en) | Breeding method of ornamental tufted ducks | |
| CN103416333A (en) | Litopenaeus vannamei base population establishing method based on genetic information and excellent properties | |
| CN107667976A (en) | A kind of producing method for seed of the high yield type reed catkins feather layer of green-shell egg of automatic sexing | |
| CN102696538A (en) | Method for breeding auto-sexing pigeon new lines | |
| CN103975898A (en) | Cultivation method for green-shelled-egg chicken pure line | |
| CN102132689A (en) | Method for breeding new species of tiejiaoma broiler chicken | |
| CN105145481A (en) | High-quality ginseng and royal jelly series chicken breeding module | |
| CN104770337A (en) | Breeding method for speckle-feather laying hens laying green-shell eggs | |
| CN104106495A (en) | Method for breeding auto-sexing pigeons by use of matching crossbreeding mode | |
| CN102100211A (en) | Method for breeding white Rex rabbit new variety | |
| CN107691354A (en) | A kind of father and mother's generation, the breeding method of the double automatic sexing broiler chicken of commercial generation | |
| CN102960271B (en) | Rotational mating breeding method for pinctada fucata groups | |
| CN103563849A (en) | Breeding method for high-yield black-skin chicken strain | |
| CN107410191A (en) | A kind of breeding method of reed catkins feather layer of green-shell egg | |
| CN101703030B (en) | The breeding method of black feather quail strain for laying eggs | |
| CN109362647A (en) | A kind of small-sized quality broiler chicken matched breeding method of low cost | |
| CN102422825B (en) | Method for establishing and breeding three matched breeding systems of Eriocheir sinensis | |
| CN107047468B (en) | Method for breeding recessive white curly eucheuma chickens | |
| CN102912017A (en) | Method for quickly and accurately identifying northern snakehead, Taiwan snakehead and hybrid snakehead |
Legal Events
| Date | Code | Title | Description |
|---|---|---|---|
| C06 | Publication | ||
| PB01 | Publication | ||
| C10 | Entry into substantive examination | ||
| SE01 | Entry into force of request for substantive examination | ||
| C14 | Grant of patent or utility model | ||
| GR01 | Patent grant | ||
| CF01 | Termination of patent right due to non-payment of annual fee |
Granted publication date: 20160106 |