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Zoysia grass plant named `Palisades`

Abstract

An asexually reproduced variety of perennial Zoysia japonica with a unique combination of characters including white stigmas, and anthers are 5.0 RP 3/6.

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A01H5/12 Leaves
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USPP11515P

United States

Inventor
Milton Charles Engelke
Current Assignee
Texas A&M University

Worldwide applications
1998 US 1999 AU JP WO 2006 JP

Application US09/078,184 events
2000-09-19
Application granted
Anticipated expiration
Expired - Lifetime
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Description

BRIEF SUMMARY OF THE INVENTION
The present invention relates to a new and distinct asexually reproduced variety of perennial zoysiagrass (Zoysia japonica (L.)) Merr.
BACKGROUND OF THE INVENTION
This invention relates to an new and distinct perennial zoysiagrass cultivar indentified as `Palisades` zoysiagrass (herein referred to as `Palisades`), that was tested as DALZ8514. `Palisades`, a Zoysia japonica, is a chance hybrid of the maternal clone `Z44`, obtained from Beltsville, MD in 1981, with an unknown pollen source from a zoysiagrass germplasm field nursery at TAES--Dallas. `Palisades` has been vegetatively propagated, and is uniform in growth expression.
For purposes of registration under the "International Convention for the Protection of New Varieties of Plants" (generally known by its French acronym as the UPOV Convention) and noting Section 1612 of the Manual of Plant Examination Procedures, the new variety of zoysiagrass of the present invention is named `Palisades` Zoysiagrass.
BRIEF DESCRIPTIONS OF THE ILLUSTRATIONS
FIG. 1 is a photograph of the leaf orientation, blades and ligule of `Palisades`.
FIG. 2 is a close-up photograph of the leaf orientation, blades and ligule of `Palisades`.
FIG. 3 is a photograph of the in florescence of `Palisades` at anthesis.
DETAILED DESCRIPTION OF THE PLANT
`Palisades` was characterized in greenhouse and field conditions. `Palisades` is a unique variety of zoysiagrass developed as described above. `Palisades` was identified as being unique and desirable and was then propagated by cutting of stolons and rhizomes, rooting them in soil, and planting of the rooted material to provide planting stock for studying performance and for comparison of morphological characters after propagation. `Palisades` has been propagated by sod, plugs, sprigs, stolons and rhizomes in greenhouse and field planting by hand and mechanical propagation in Dallas, Tex. and is uniform in growth expression. Seed reproduction with self-fertility is not common in the Zoysia sp. No seedling establishment from `Palisades` has been noticed in either greenhouse or field studies.
`Palisades` is asexually propagated by stolons and rhizomes. It has an intermediate to rapid establishment rate. When spring (stolons) or plug plantings are made in early May, the area should achieve 60-70% coverage in Dallas, Tex. in 90-120 days, with complete coverage in 180-200 days, in comparison to `Meyer` zoysiagrass which will require 360-400 days for full coverage. The stolons of `Palisades` have a mean internode length of 29.9 mm between the second and third nodes, with a mean internode width of 1.0 mm and node diameter of 1.7 mm (Table 1). The stolons of `Palisades` root adventitiously at the nodes. Color notations of plant tissues were based on the Munsell Color Charts for Plant Tissues, Munsell Color, Baltimore, Md., 1977. Light quality, photoperiod, and general growth of the plants affect color notations. The internode stolon color of `Palisades` stolons exposed to full sun is 5R 3/4.
Leaf blades of `Palisades` are rolled in the bud, and are flat and stiff. The mean leaf blade length of `Palisades` is 75.6 mm and average width is 3.4 mm (Table 2). The leaf sheath of `Palisades` is distinct from `Meyer`(unpatented) and `Crowne` (U.S. Plant Patent application Ser. No. 09/076,973 at a length of 51.2 mm (Table 3). The hairs on the abaxial leaf surface of `Palisades` are sparse in number and are approximately 0.9 mm long. Measured under greenhouse conditions in January 1996, the genetic, adaxial leaf color of `Palisades` is 2.5G 5/6 with `El Toro` (U.S. Plant Pat. No. 5845) having a leaf color of 2.5GY 5/2, and `Meyer` having a color of 2.5G 3/4. The ligule of `Palisades` is a fringe of silky hairs, approximately 2.8 mm in length for the longest hairs.
`Palisades` has 5RP 3/6 colored anthers and white colored stigmas, undistinguished in shade of color. The inflorescence of `Palisades` is a terminal spike-like raceme, with spikelets on short pedicels. `Palisades` has a mean length floral region of 27.2 mm, with a mean of 27.8 florets per raceme.
Thatch development is a function of age of stand, types of soils and other environments and cultural conditions. Thatch development was determined in 3 year old field plantings comparing eight commercially available cultivars. `Emerald`, `Miyako` (U.S. Plant Pat. No. 10,187) and `Zeon` (unpatented) produced the greatest depth of thatch (26.3 mm); `Crowne`, `Palisades` and `DeAnza` produced an intermediate amount of thatch (25,23 and 23 mm, respectively), and Cavalier and Meyer produced the least amount of thatch (20 and 19 mm respectively).
The chromosome number of `Palisades` is 40.
`Palisades` is rated as having low supplemental water needs similar to `Crowne` and `El Toro` to maintain active growing turf (Table 4). When `Palisades` was compared with 59 other zoysiagrasses for salinity tolerance, it ranked number 34 in performance and was superior to `Meyer` (Table 5). When compared to other commercial varieties for root growth, `Palisades` was not different from `El Toro` or `Meyer` in average root depth, root weight, or clipping weights (Table 6).
`Palisades` was entered in the 1991 National Turfgrass Evaluation Program (NTEP). `Palisades`, in the 1991 National Turfgrass Evaluation Program Zoysiagrass Trial for the years 1992-1995 had mean quality ratings of 5.8, 5.8, 5.5 and 5.4 for overall quality (Table 7).
`Palisades` rated among the best for the broad leaf textural class 4. `Palisades` shows intermediate resistance to the zoysiagrass mite (Table 8). The varieties `Meyer`, `Belair` (unpatented), and most experimental zoysiagrasses are very susceptible to the mite. This mite has been found in Maryland, Texas, and Florida, all extensive zones of use for zoysiagrasses.
`Palisades` is distinguished from other zoysiagrasses by its combined unique morphological differences in leaf width, length, and floral characters of sheath and seed head length. It is a medium-coarse textured, thick stolon, and rhizome producing grass with good recuperative ability. It has good shade tolerance and low water use needs. It is moderate in salinity tolerance. `Palisades` is intermediate to superior in its average growth rate and has good to excellent winter hardiness and will persist in regions north to Kansas, Missouri and Illinois.
              TABLE 1                                                     
______________________________________                                    
Internode length as measured between the second and third nodes,          
internode diameter of the third internode, and node diameter of the       
third node measured on zoysiagrass plants. Plants were growing in a       
growth chamber with a 14-hr daylength, March 1995                         
            Internode   Internode                                         
                                 Node                                     
            length      diameter diameter                                 
Genotype    mm          mm       mm                                       
______________________________________                                    
`Crowne`    17.7a*      0.9abc   1.1 bc                                   
`Cavalier`  18.4a       1.4abc   1.4abc                                   
(U.S. Plant                                                               
Pat. No. 10,778)                                                          
`Meyer`     24.1a       2.0a     2.0a                                     
`Palisades` 29.9a       1.0a     1.7ab                                    
`El Toro`   39.98a      1.4abc   1.4abc                                   
______________________________________                                    
 *Analysis of variance by General Linear Models, with means followed by th
 same letter not significantly different using Tukey's Studentized Range  
 (HSD), alpha = 0.05. Only selected means presented.                      
              TABLE 2                                                     
______________________________________                                    
Leaf blade width and length measured on the third youngest                
leaf of zoysiagrass. Plants were growing in a growth chamber with a       
14-hr daylength, March 1995                                               
              Blade width                                                 
                        Blade length                                      
Genotype      mm        mm                                                
______________________________________                                    
`Cavalier`    1.3d*     60.7cd                                            
`Meyer`       2.6 c     74.3 bc                                           
`Crowne`      3.0abc    72.7 bc                                           
`El Toro`     3.3abc    68.3 bcd                                          
`Palisades`   3.4ab     75.6 bc                                           
______________________________________                                    
 * Analysis of variance by General Linear Models, with means followed by  
 the same letter not significantly different using Tukey's Studentized    
 Range (HSD), alpha = 0.05. Only selected means presented.                
              TABLE 3                                                     
______________________________________                                    
Zoysiagrass leaf sheath length measurements taken                         
February 1988 on the fourth youngest leaf from greenhouse                 
grown plants                                                              
                 Sheath length                                            
Genotype         mm                                                       
______________________________________                                    
`Emerald`(unpatented)                                                     
                 16.3 ij                                                  
`Cavalier`       28.9 ef                                                  
`Meyer`          30.7 cd                                                  
`Crowne`         42.9 b*                                                  
`Palisades`      51.25a                                                   
______________________________________                                    
 *Means followed by the same letter are not significantly different using 
 the WallerDuncan k ratio test (k ratio = 100). Only selected means       
 presented.                                                               
              TABLE 4                                                     
______________________________________                                    
Supplemental irrigation water requirement for commercial and              
experimental zoysiagrasses during July 1989 through August 1991 at        
Dallas, Texas*                                                            
                   Irrigation                                             
                   Requirement                                            
              Textural                                                    
                     1989   1990   1991                                   
              Class  mm/yr        Mean                                    
______________________________________                                    
`Diamond`       1        461    435  567  488                             
(U.S. Plant                                                               
Pat. No. 10,636)                                                          
DALZ8501        1        449    544  429  474                             
FC13521         3        482    448  443  457                             
DALZ8517        3        475    402  487  455                             
`Emerald`       3        464    343  503  437                             
DALZ8506        3        458    379  455  431                             
DALZ8515        3        469    419  394  427                             
DALZ8508        2        447    379  398  408                             
DALZ8510        3        449    310  413  390                             
`Cashmere`      1        435    424  311  390                             
U.S. Plant                                                                
Pat. No. 6,529                                                            
`Cavalier`      3        464    175  441  360                             
DALZ8504        2        478    363  138  326                             
DALZ8503        2        441    280  193  305                             
DALZ8511        2        451    353  200  304                             
DALZ8516        2        462    377  25   288                             
`Meyer`         2        450    321  74   276                             
`Korean Common`(unpatented)                                               
                4        470    174  88   244                             
`El Toro`       4        417    21   6    148                             
`Palisades`     4        358    26   12   132                             
`Crowne`        4        256    12   12   93                              
                MSD.sup.†                                          
                         129    169  242  155                             
                Rainfall.sup.‡                                 
                         1092   1118 1143 1118                            
______________________________________                                    
 .sup.†MSD, minimum significant difference for comparison of means 
 within columns based on the WallerDuncan kratio ttest where k = 100.     
 .sup.‡Total annual precipitation in mm.                       
 *White et al. 1993.                                                      
              TABLE 5                                                     
______________________________________                                    
* Average percent shoot salt injury (average of 20 rating dates) on       
zoysiagrass entries in the 1991 NTEP.sup.† Trials                  
Entry         Source    Species.sup.‡                          
                                  % Injury                                
______________________________________                                    
`Diamond`     NTEP 20   matrella  33ab.sup..English Pound.                
`El Toro`     NTEP 13   japonica  38a-e                                   
`Emerald`     NTEP 10   jap x tenu                                        
                                  42a-h                                   
`Cavalier`    NTEP 17   matrella  42a-h                                   
`Crowne`      NTEP 15   japonica  46 e-k                                  
`Palisades`   NTEP 14   japonica  46 e-k                                  
`Belair`      NTEP 11   japonica  50 f-k                                  
`Meyer`       NTEP 09   japonica  58 lmn                                  
`Korean Common`                                                           
              NTEP 07   japonica  76 pq                                   
______________________________________                                    
 * Selected data set; complete data set includes 59 varieties and         
 cultivars. .sup.† NTEP = National Turfgrass Evaluation Program.   
 .sup.‡ Species identity. .sup..English Pound. Means followed b
 the same letter are not significantly different, based on the WallerDunca
 kratio ttest (kratio = 100)                                              
 In Marcum, K. B., M. C. Engelke, S. J. Morton and C. Dayton. 1994.       
 Salinity tolerances of selected bermudagrass an zoysiagrass genotypes. TX
 Turfgrass Res. 1993, Consolidated Prog. Rep. PR 5140: 105-107.           
              TABLE 6                                                     
______________________________________                                    
Average mean root depth of zoysiagrass grown in flexible tubes in         
greenhouse studies, Dallas, TX                                            
         Average Mean  Total Root                                         
                                Clipping                                  
         Root Depth    Weight   Weights                                   
Variety  mm            mg       mg                                        
______________________________________                                    
`Cavalier`                                                                
         255           278      243                                       
`Belair` 296           330      286                                       
`Palisades`                                                               
         318           457      452                                       
`Emerald`                                                                 
         330           461      241                                       
`Meyer`  333           411      466                                       
`Crowne` 355           497      406                                       
`El Toro`                                                                 
         356           473      391                                       
MSD      79            161      267                                       
______________________________________                                    
 *MSD = minimum significant difference for comparison of means within     
 columns based on the WallerDuncan kratio test where k = 100.             
 In Marcum, K. B., M. C. Engelke, S. J. Morton, and R. H. White. 1995.    
 Rooting characteristics and associated drought resistance of             
 zoysiagrasses. Agron. J. 87:534-538.                                     
              TABLE 7                                                     
______________________________________                                    
Mean Turfgrass quality ratings of vegetative zoysiagrass cultivars grown  
in the National Turfgrass Evaluation Program at 23 locations in the U.S.  
for 1992, 1993, 1994, and 1995                                            
            Leaf                                                          
            Texture                       Overall                         
Variety     Class   1992   1993 1994 1995 4-Yr Avg.                       
______________________________________                                    
`Cavalier`  .sup. 3.sup.1                                                 
                    6.0    6.2  5.9  6.0  5.9                             
TC2033      2       5.8    6.1  6.1  6.0  5.9                             
`Sunburst`(unpatented)                                                    
            2       5.8    5.9  5.8  5.9  5.8                             
TC 5018             5.8    5.8  5.9  5.7  5.8                             
`Emerald`   2       5.7    6.2  6.0  5.7  5.8                             
`Omni`(unpatented)                                                        
            2       5.6    6.1  6.1  6.0  5.7                             
QT 2004             5.6    6.0  5.9  5.6  5.6                             
DALZ8508    2       5.6    6.1  5.7  5.6  5.6                             
`Palisades` 4       5.8    5.8  5.5  5.4  5.6                             
DALZ9006    3       5.6    6.0  5.6  5.5  5.6                             
`Crowne`    4       5.8    5.8  5.5  5.4  5.6                             
`El Toro`   4       5.8    5.6  5.3  5.4  5.5                             
CD 259-13           5.3    5.5  5.7  5.5  5.4                             
`Meyer`     2       5.3    5.7  5.8  5.5  5.4                             
QT 2047             5.4    5.4  5.3  5.2  5.3                             
`Belair`    2       5.0    5.6  5.6  5.0  5.2                             
DALZ 8516   2       4.7    5.4  5.0  5.0  4.9                             
`Diamond`   1       4.4    5.0  4.6  4.4  4.4                             
DALZ8501    1       4.9    4.3  4.0  4.0  4.3                             
DALZ8701    1       4.2    4.1  3.7  3.6  3.8                             
LSD VALUE           0.2    0.2  0.2  0.2  0.2                             
______________________________________                                    
 To determine statistical differences among entries, subtract one entry's 
 mean from another entry's mean. Statistical difference occurs when this  
 value is larger than the corresponding LSD value (LSD 0.05). .sup.1      
 Textural class of zoysiagrass where 1 = short, narrow leaves; 2 = short, 
 wide leaves; 3 = long, narrow leave; and 4 = long, wide leaves.          
 In National Zoysiagrass Test  1991. Final Report 1992-95, NTEP No. 9615; 
 (Table 4); United States Department of Agriculture, Agricultural Research
 Service, Beltsville Agricultural Research Center, Beltsville, MD 20705.  
              TABLE 8                                                     
______________________________________                                    
Mean number of mite-damaged leaves per zoysiagrass plant (N = 18)         
                                  Combined                                
Cultivar  April 3, 1992                                                   
                       April 24, 1992                                     
                                  Dates                                   
______________________________________                                    
DALZ9006  0.2a*        0.2a       0.2a                                    
DALZ8508  0.7b         0.6ab      0.6b                                    
DALZ8516  1.8d         0.3ab      1.0b                                    
`Emerald` 1.3bc        0.9b       1.1b                                    
DALZ8501  1.4cd        2.6c       2.0c                                    
`Crowne`  5.4ef        2.3c       3.9d                                    
`El Toro` 5.6ef        2.8c       4.2d                                    
TC2033    4.4e         4.3d       4.4d                                    
`Palisades`                                                               
          7.5fg        4.0cd      5.8e                                    
CD2031    7.6gh        7.9e       7.8f                                    
Diamond   7.8gh        9.4ef      8.6g                                    
DALZ8701  9.3h         9.4ef      9.4g                                    
`Cavalier`                                                                
          9.1h         9.8ef      9.5g                                    
`Meyer`   9.9h         9.9ef      9.9g                                    
`Belair`  10.0h        9.9f       9.9g                                    
JZ-1      10.0h        9.9f       10.0g                                   
______________________________________                                    
 *Data transformed using log (X + 0.5) for analysis. Means in a column    
 followed by the same letter are not significantly different by           
 WallerDuncan kratio test (k = 100) (P = 0.05).                           
 Reinert, J. A., M. C. Engelke, and S. J. Morton. 1993. Zoysiagrass       
 resistance to the zoysiagrass mite, Eriphyes zoysiae (Acari: Eriophyidae)
 In R. N. Carrow, N. E. Christians, and R. C. Shermans (Eds.).            
 International Turfgrass Society Research Journal 7. Intertec Publishing  
 Corp., Overland Park, KS. pp. 349-352.                                   

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I claim:
1. A new and distinct cultivar of Zoysia japonica plant as herein shown and described.