CN106171745B - A kind of method of measurement function group seedling Niche Differentiation on light resource axis - Google Patents
A kind of method of measurement function group seedling Niche Differentiation on light resource axis Download PDFInfo
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- CN106171745B CN106171745B CN201610550337.9A CN201610550337A CN106171745B CN 106171745 B CN106171745 B CN 106171745B CN 201610550337 A CN201610550337 A CN 201610550337A CN 106171745 B CN106171745 B CN 106171745B
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- A—HUMAN NECESSITIES
- A01—AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
- A01G—HORTICULTURE; CULTIVATION OF VEGETABLES, FLOWERS, RICE, FRUIT, VINES, HOPS OR SEAWEED; FORESTRY; WATERING
- A01G17/00—Cultivation of hops, vines, fruit trees, or like trees
- A01G17/005—Cultivation methods
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- A—HUMAN NECESSITIES
- A01—AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
- A01G—HORTICULTURE; CULTIVATION OF VEGETABLES, FLOWERS, RICE, FRUIT, VINES, HOPS OR SEAWEED; FORESTRY; WATERING
- A01G23/00—Forestry
Abstract
The invention discloses a kind of methods of measurement function group seedling Niche Differentiation on light resource axis, comprising the following steps: field investigation, with choosing sample；Equal sample, divide plaque type；The division of functional group is carried out according to the light of different seedling；Measure seedling corresponding index in field natural habitat each typical patch stage；It measures seedling and simulates corresponding index in each stage in manual light control；Measure the morphosis and physiological and biochemical index of seedling；The drafting of different function group seedling ecological bitmap on light resource axis.The present invention provides an important method for the quantization of Niche Differentiation, to explain that forest community Wuling mountain mechanism provides an important evidence, brings important realistic meaning for the quantitative study of plant population ecology.
Description
Technical field
It is ecological on light resource axis that the present invention relates to measuring method technical fields more particularly to a kind of measurement function group seedling
The method of position differentiation.
Background technique
Ecological niche (ecological niche) refers to a population in the ecosystem, occupied on time and space
Position and its functional relationship and effect between related population.The concept of ecological niche is applied first by Grinnell earliest
, he thinks that there are two the species established for a long time to have duplicate ecological niche in a fauna.Later Elton is mentioned
" relationship that ecological niche is status and it and things and natural enemy of the species in its living environment " out, contains Trophic Niche
The meaning.
In natural environment, each specific position has different types of biology, movable and with other biologies
Relationship depends on its special construction, physiology and behavior, therefore with the unique ecological position of oneself.As each biology occupies respectively
From space, there is respective function and nutrition position in group, and in environmental changes such as temperature, humidity, soil, illumination
The status occupied in gradient.In areal, biological type is abundanter, between species in order to nutrition, living space or other
Resource and the competition occurred is fiercer, in this way, the realized niche occupied to a certain particular species may be smaller and smaller.
As a result, during evolution, in two ecologies very close to species towards occupying different space (habitat differentiation), eat
Break up on different things (specialization in feeding habits), different activity time (time differentiation) or other ecological habits, it is competing to reduce
The tensity striven, to make to be likely to form balance between two kinds and coexist.
Niche Differentiation has been the widely used term of domestic and international field of ecology and research hotspot.Due to species diversity
The principles such as property, Species Competition, the succession of community, Evolution of Population are all closely related with Niche Differentiation theory, therefore Niche Differentiation
Theory gradually receives the extensive attention of domestic and foreign scholars.
So far, Niche Differentiation problem is always the focus of ecological study, but as described above, and many researchs are all
It is that the measuring method of suitable quantization Niche Differentiation is lacked based on theory.Therefore, it is necessary to which specific aim proposes a kind of new side
Method improves correlative study.
Summary of the invention
Technical problem to be solved by the present invention lies in providing, a kind of measurement function group seedling is ecological on light resource axis
The method of position differentiation.
In order to solve the above technical problems, the present invention is provided based on patch pattern scale measurement function group seedling in light resource axis
The method of upper Niche Differentiation, comprising the following steps:
(1): field investigation
With choosing sample；
(2): plaque type divides
Each sample is divided into multiple patches, and determines the type in different patch stage: gap patches, establish patch,
Mature patch and decline patch；
(3): the division of different tree species seedling functional group
The four-stage of forest cycle includes: woods window stage, establishment stage, the stage of ripeness and decline stage；
The frequency that the four-stage different tree species of forest cycle occur by inquiry, in conjunction with its light property, by different tree species
Seedling is divided into pioneer's functional group, transition stage functional group and the top functional group three classes of resistance to shade；
(4): field nature simulation
The Potted orchard for choosing same size moves to each typical patch stage, and seedling is to different patch under simulation natural habitat
The adaptation and response of interior intensity of illumination measure the variation that seedling corresponding index is respectively handled in the different patch stage；
(5): manual light control's simulation
Setting control light experiment, the raw seedling of 1-3 of transplanting nature growth, simulating forest patch are developed naturally to next rank
Section moves on to the lighting process under next patch gradient, measures corresponding index after controlling light processing 5-15d；Simulating forest spot
It is interfered to form woods window in block nature growth course, is i.e. after control light processing 5-15d, moves on under the patch gradient in woods window stage
Lighting process measures corresponding index；
(6): morphosis and Determination of Physiological And Biochemical Indices
1. the measurement of morphosis
Anatomical structure, the measurement for constructing behavior and spatial orientation index；
2. the measurement of physiological and biochemical index:
Photosynthetic production capacity, the level of resources utilization and photosensitizing substance conversion ratio measurement；
(7): the differentiation of different function group seedling ecological niche on light resource axis
Using the intensity of illumination of different phase in forest cycle function dynamic as abscissa, respectively with different function group seedling
Morphosis index and physiological and biochemical index are ordinate, draw ecological bitmap of the different function group seedling on light resource axis.
Preferably, manual light control's simulated experiment black shading screen shades, and shade frame uses triangle frame, artificial to hide
4 light gradients are selected in shade, simulate woods window stage, establishment stage, the intensity of illumination in the stage of ripeness and decline stage respectively；By cool canopy
North orientation open as channel, remaining each Xiang Junyong shading screen covering；By the middle part for being located proximate to cool canopy of Potted orchard.
Preferably, the measuring method of the anatomical structure index includes: to randomly select seedling from every group of light processing, by not
It is equidirectional to take the good current-year branch of growth and development in the middle part of tree crown, it is taken back in laboratory with ice chest；Select moderate fresh of thickness
Several pieces of leaf, the middle part of every leaf is cut, FAA fixer is fixed, and paraffin sections are made, and analyzes microstructure；External application out of office 3% penta
Dialdehyde fixer is fixed, and is taken back laboratory and is transferred to fixation in 2% osmic acid fixer, ultra-thin section is made, analyzes ultra microstructure.
Preferably, the microstructure include: leaf overall thickness, the upper and lower epidermal thickness of leaf, cutin film thickness, epicuticle with
Endothelium interlayer mesophyll tissue thickness, lower epidermis and endothelium interlayer mesophyll tissue thickness, the thickness of endothelium confluent monolayer cells, transfusion tissue with
Vascular bundle thickness and the diameter of tracheid etc..
Preferably, the ultra microstructure includes: chloroplaset morphosis, the distribution of chloroplaset and thylakoid shape etc..
It is preferably, described that construct behavior and the measuring method of spatial orientation index include: when carrying out seedling parsing, from tree
The tip starts to look into several blade quantities by side shoot downwards, measures increment, measures branch angle with graphometer, and measure each side shoot in trunk
On distribution, the long, crotch angle of branch etc.；After root system is all dug out, root range, root depth are first measured, several lateral root numbers is looked into, then will
All lateral roots are cut from base portion, measure root long and root diameter, calculation side root surface area respectively.
Preferably, the measuring method of the photosynthetic production capacity index includes: by the dark adaptation in leaf presss from both sides of the blade of selection
15-25min measures corresponding fluorescence parameter with portable modulation luminoscope: initial fluorescence, maximum fluorescence, temperture of leaves, it is photosynthetic effectively
Radiation, II maximum quantum yield of PS, II photo-quantum efficiency of PS, Relative electron transport rate, Photochemical quenching coefficient and non-photochemical
Learn quenching coefficient；Net Photosynthetic Rate, transpiration rate, intercellular CO are measured with photosynthetic apparatus2Concentration, stomatal conductance；Choose every group
The current year leave piece of the lower 3 plants of seedling of light processing, is taken back in laboratory with ice chest, every plant of blade is shredded mixing, uses ultraviolet spectrometry
Photometric determination chlorophyll a, b and carotenoid content.
Preferably, the measuring method of the level of resources utilization index includes: seedling leaves sample that will fetch in 70-90
DEG C drying after pulverize and sieve, using the full nitrogen of Kjeldahl nitrogen determination；Photosynthetic had according to net primary production with what is absorbed
Amount of radiation ratio is imitated to calculate light utilization ratio, net photosynthetic rate calculates carbon utilization efficiency with Dark respiration rate ratio,
The product of Nitrogen productivity and nitrogen mean residence time calculates nitrogen use efficiency.
Preferably, the measuring method of the photosensitizing substance conversion ratio index includes: with Bomass allocation and blade assimilation effect
Rate reacts photosensitizing substance conversion ratio, several plants of seedling are randomly selected in every group of light processing, claims each organ to fresh weight respectively, and
Sampling take back in laboratory, 70-90 DEG C drying 36-60 hour, title root, stem, leaf each section organ dry weight, conversion dry matter
Amount, and calculate following index: root compares again, stem compares again, leaf mass ratio, ground/Underground biomass, Gao Jingchong when relative growth
Rate etc.；In order to inquire into the assimilatory efficiency difference of each level blade of tree crown, weigh respectively to each sequence side shoot, and will work as
Year raw branch individually samples weighing, with each organic unit's time biomass total amount in non-assimilation part, current year Leaf biomass and a
Based on the total leaf area of body, the gross assimilation efficiency of blade is calculated, then calculates each organ according to Bomass allocation ratio
Blade assimilatory efficiency.
It preferably, is the sample north subtropical Da Bie Mountain area Mixed Evergreen-deciduous Broad-leaved Forests.
The present invention has the advantages that the present invention is by providing one kind based on patch pattern scale measurement function group seedling in light
The method of Niche Differentiation is plant species to explain that forest community Wuling mountain mechanism provides an important evidence on resource axis
The ecological quantitative study of group brings important realistic meaning.
Detailed description of the invention
Fig. 1 is the differentiation figure of different function group seedling provided in an embodiment of the present invention ecological niche on light resource axis.
Specific embodiment
It elaborates below to the embodiment of the present invention, the present embodiment carries out under the premise of the technical scheme of the present invention
Implement, the detailed implementation method and specific operation process are given, but protection scope of the present invention is not limited to following implementation
Example.
The present embodiment includes the following steps:
(1): field investigation
Choose north subtropical Da Bie Mountain area Mixed Evergreen-deciduous Broad-leaved Forests: Cyclobalanopsis gracilis-short handle Serrata oakery, leaflet are green
Ridge-Mao Lilin, Qinggang oak-oriental white oak-yellow wingceltis woods and cork oak-cyclobalanopsis gleuca, with respectively selecting the sample of 4 100m × 100m；
(2): plaque type divides
Each sample is divided into the patch of 400 5m × 5m, the type in different patch stage is determined according to table 1；
1 different patch stage type of table calibrates standard really
(3): the division of different tree species seedling functional group
Demand according to the growth of seedling to light resource is classified as pioneer's functional group, transition stage functional group and resistance to shady top
Grade functional group.Specific judgment criteria is that the seedling of pioneer's functional group can only be not present in closing in woods window and open ground discovery
Hayashishita；Rather than the seedling of pioneer's functional group can exist under complete stand, seldom occur in woods window.Non- pioneer's functional group again may be used
To be divided into transition stage functional group and the resistance to top functional group of shade.Wherein, the resistance to top functional group of shade has intermediate light demand,
Although can survive in hayashishita, need higher luminous environment that can just grow up to bigger individual.Forest cycle by inquiry
The frequency that four-stage (woods window stage, establishment stage, the stage of ripeness and decline stage) different tree species occur, in conjunction with its light
Property, by the different tree species seedling grown naturally in Mixed Evergreen-deciduous Broad-leaved Forests (Cyclobalanopsis gracilis, short handle Serrata oak, Mao Li, Qinggang
Oak, oriental white oak, yellow wingceltis and cork oak) pioneer's functional group, transition stage functional group and the top functional group three classes of resistance to shade are divided, such as table 2
It is shown；
The division of the different plant functional groups of table 2
(4): field nature simulation
The Potted orchard (1 plant of seedling of each Potted orchard) for choosing same size moves on to each typical case's patch stage under natural habitat
Middle part, simulate natural habitat under adaptation and response of the seedling to intensity of illumination in different patch, measurement the woods window stage, establish rank
The variation of seedling corresponding index is respectively handled in section, the stage of ripeness and decline stage；(5): manual light control's simulation
Manual light control's simulated experiment black shading screen shades.4 light gradients are selected in artificial shade, simulate woods window rank respectively
Section, establishment stage, the intensity of illumination in the stage of ripeness and decline stage.It, will for the sake of convenient for normal management and periodic measurement convenience
The north orientation of cool canopy is opened as channel, remaining each Xiang Junyong shading screen covering.In order to avoid shading each other between cool canopy to young in canopy
Seedling has an impact, as far as possible by the middle part for being located proximate to cool canopy of Potted orchard.
Setting control light experiment, the Cyclobalanopsis gracilis of Mixed Evergreen-deciduous Broad-leaved Forests hayashishita of transplanting nature growth, short handle Serrata oak,
Seven kinds of Mao Li, Qinggang oak, oriental white oak, yellow wingceltis, cork oak raw seedling of 1-3 use 200 basin Potted orchards under each lighting gradients, hide
Shady frame uses triangle frame.
Simulating forest patch was developed naturally to next stage, that is, after controlling light processing 10d, was moved on under next patch gradient
Lighting process, measure corresponding index.
It is interfered to form woods window in simulating forest patch nature growth course, is i.e. after control light processing 10d, moves on to woods window rank
Lighting process under the patch gradient of section, measures corresponding index；
(6): morphosis and Determination of Physiological And Biochemical Indices
1. the measurement of morphosis
The measurement of anatomical structure index: randomly selecting 5 plants of seedling from every group of light processing, and every plant takes growth hair in the middle part of tree crown
Good current-year branch 4 (taking by different directions, to eliminate difference caused by the difference of direction) is educated, takes back laboratory with ice chest
It is interior.By downward the 3rd Shu Yezhong of number of shoot apex, 5 pieces of fresh leaf that thickness is moderate are selected, cut the middle part of every leaf, FAA fixer is solid
It is fixed, take back laboratory and be made paraffin sections, analysis microstructure (the upper and lower epidermal thickness of leaf overall thickness, leaf, cutin film thickness, on
Epidermis and endothelium interlayer mesophyll tissue's thickness and lower epidermis and endothelium interlayer mesophyll tissue thickness, endothelium confluent monolayer cells thickness, turn
The indexs such as the diameter of defeated tissue and vascular bundle thickness and tracheid)；3% glutaraldehyde fixer of external application out of office is fixed, and laboratory is taken back
It is transferred in 2% osmic acid fixer and fixes, ultra-thin section is made, analyze the ultra microstructure (distribution of chloroplaset morphosis, chloroplaset
With thylakoid shape etc.).
It constructs the measurement of behavior and spatial orientation index: when carrying out seedling parsing, being looked by side shoot downwards since treetop
Number blade quantities, measure increment, measure branch angle with graphometer, and measure that distribution of each side shoot on trunk, branch be long, branch
Angle etc..After root system is all dug out, root range, root depth are first measured, several lateral root numbers is looked into, then cuts all lateral roots from base portion
Under, root long and root diameter, calculation side root surface area are measured respectively.
2. the measurement of physiological and biochemical index:
Photosynthetic production capacity: the blade of selection is pressed from both sides into interior dark adaptation 20min in leaf, measures phase with portable modulation luminoscope
The fluorescence parameter answered: initial fluorescence, maximum fluorescence, temperture of leaves, photosynthetically active radiation, II maximum quantum yield of PS, II light quantum of PS
Efficiency, Relative electron transport rate, Photochemical quenching coefficient and non-Photochemical quenching coefficient.Net photosynthesis is measured with photosynthetic apparatus
Rate, transpiration rate, intercellular CO2Concentration, stomatal conductance.The current year leave piece for choosing the lower 3 plants of seedling of every group of light processing, uses ice chest
It takes back in laboratory, every plant of blade is shredded into mixing, measure chlorophyll a, b with ultraviolet specrophotometer and carotenoid contains
Amount.
The level of resources utilization: the seedling leaves sample fetched is pulverized and sieved after 80 DEG C of drying, is surveyed using conventional method
Fixed full nitrogen (Kjeldahl's method).Light benefit is calculated according to net primary production and the photosynthetically active radiation amount ratio absorbed
With efficiency, net photosynthetic rate calculates carbon utilization efficiency with Dark respiration rate ratio, and Nitrogen productivity is average stagnant with nitrogen
The product of time is stayed to calculate nitrogen use efficiency.
Photosensitizing substance conversion ratio: photosensitizing substance conversion ratio is reacted with Bomass allocation and blade assimilatory efficiency.At every group
5 plants of seedling are randomly selected in light processing, claim each organ to fresh weight respectively, and sample and take back in laboratory, it is small in 80 DEG C of drying 48
When, claim root, stem, leaf each section organ dry weight, convert amount of dry matter.And calculate following index: root is again than (root weight/total biology
Amount), stem compares than (stem weight/total biomass), leaf mass ratio (Ye Chong/total biomass), ground/Underground biomass, high stem again again
(plant height/stem biomass) and relative growth rate etc..In order to inquire into the assimilatory efficiency difference of each level blade of tree crown, respectively to each
A sequence side shoot is weighed, and current-year branch is individually sampled to weighing.With each organic unit's time biology in non-assimilation part
Based on measuring total amount, current year Leaf biomass and the total leaf area of individual, the gross assimilation efficiency of blade is calculated, then according to biology
Amount allocation proportion calculates the blade assimilatory efficiency of each organ；
(7): the differentiation of different function group seedling ecological niche on light resource axis
Intensity of illumination with forest cycle function dynamic middle forest window stage, establishment stage, the stage of ripeness and decline stage is
Abscissa, respectively with the morphosis index (anatomical structure constructs behavior and root structure) and physiology of different function group seedling
Biochemical indicator (photosynthetic production capacity, the level of resources utilization and photosensitizing substance conversion ratio) is ordinate, draws different function group children
Ecological bitmap (Fig. 1) on Miao Guang resource axis.
The foregoing is merely illustrative of the preferred embodiments of the present invention, is not intended to limit the invention, all in essence of the invention
Made any modifications, equivalent replacements, and improvements etc., should all be included in the protection scope of the present invention within mind and principle.
Claims (10)
1. a kind of method of measurement function group seedling Niche Differentiation on light resource axis, which comprises the steps of:
(1): field investigation
With choosing sample；
(2): plaque type divides
Each sample is divided into multiple patches, and determines the type in different patch stage: gap patches establish patch, maturation
Patch and decline patch；
(3): the division of different tree species seedling functional group
The four-stage of forest cycle includes: woods window stage, establishment stage, the stage of ripeness and decline stage；
The frequency that the four-stage different tree species of forest cycle occur by inquiry, in conjunction with its light property, by different tree species seedling
It is divided into pioneer's functional group, transition stage functional group and the top functional group three classes of resistance to shade；
(4): field nature simulation
The Potted orchard for choosing same size moves to each typical patch stage, and seedling is to light in different patch under simulation natural habitat
According to the adaptation and response of intensity, the variation that seedling corresponding index is respectively handled in the different patch stage is measured；
(5): manual light control's simulation
Setting control light experiment, the raw seedling of 1-3 of transplanting nature growth, simulating forest patch were developed naturally to next stage,
After controlling light processing 5-15d, the lighting process under next patch gradient is moved on to, corresponding index is measured；Simulating forest patch
It is interfered to form woods window in natural growth course, is i.e. after control light processing 5-15d, moves on to the light under the patch gradient in woods window stage
According to processing, corresponding index is measured；
(6): morphosis and Determination of Physiological And Biochemical Indices
1. the measurement of morphosis
Anatomical structure, the measurement for constructing behavior and spatial orientation index；
2. the measurement of physiological and biochemical index:
Photosynthetic production capacity, the level of resources utilization and photosensitizing substance conversion ratio measurement；
(7): the differentiation of different function group seedling ecological niche on light resource axis
Using the intensity of illumination of different phase in forest cycle function dynamic as abscissa, respectively in the form of different function group seedling
Structure index and physiological and biochemical index are ordinate, draw ecological bitmap of the different function group seedling on light resource axis.
2. a kind of method of measurement function group seedling Niche Differentiation on light resource axis according to claim 1, special
Sign is that manual light control's simulated experiment black shading screen shades, and shade frame uses triangle frame, and 4 light are selected in artificial shade
Gradient simulates woods window stage, establishment stage, the intensity of illumination in the stage of ripeness and decline stage respectively；The north orientation of cool canopy is open-minded
As channel, remaining each Xiang Junyong shading screen covering；By the middle part for being located proximate to cool canopy of Potted orchard.
3. a kind of method of measurement function group seedling Niche Differentiation on light resource axis according to claim 1, special
Sign is that the measuring method of anatomical structure index includes: to randomly select seedling from every group of light processing, takes tree crown by different directions
The good current-year branch of middle part growth and development, is taken back in laboratory with ice chest；It several pieces of the fresh leaf for selecting thickness moderate, cuts
The middle part of every leaf, FAA fixer are fixed, and paraffin section is made, and analyze microstructure；3% glutaraldehyde fixer of external application out of office is solid
Calmly, it takes back laboratory and is transferred to fixation in 2% osmic acid fixer, ultra-thin section is made, analyzes ultra microstructure.
4. a kind of method of measurement function group seedling Niche Differentiation on light resource axis according to claim 3, special
Sign is that microstructure includes: leaf overall thickness, the upper and lower epidermal thickness of leaf, cutin film thickness, epicuticle and endothelium interlayer mesophyll
Tissue thickness, lower epidermis and endothelium interlayer mesophyll tissue thickness, the thickness of endothelium confluent monolayer cells, transfusion tissue and vascular bundle thickness and
The diameter of tracheid.
5. a kind of method of measurement function group seedling Niche Differentiation on light resource axis according to claim 3, special
Sign is that ultra microstructure includes: chloroplaset morphosis, the distribution of chloroplaset and thylakoid shape.
6. a kind of method of measurement function group seedling Niche Differentiation on light resource axis according to claim 1, special
Sign is, construct behavior and spatial orientation index measuring method include: when carrying out seedling parsing, since treetop downwards by
Side shoot looks into several blade quantities, measures increment, measures branch angle with graphometer, and measures distribution of each side shoot on trunk, branch
Long, crotch angle；After root system is all dug out, it is deep first to measure root range, root, looks into several lateral root numbers, then by all lateral roots from base
Portion is cut, and measures root long and root diameter, calculation side root surface area respectively.
7. a kind of method of measurement function group seedling Niche Differentiation on light resource axis according to claim 1, special
Sign is that the measuring method of photosynthetic production capacity index includes: that the blade of selection is pressed from both sides interior dark adaptation 15-25min in leaf, with just
Take formula modulation luminoscope and measure corresponding fluorescence parameter: initial fluorescence, maximum fluorescence, temperture of leaves, photosynthetically active radiation, PS II are maximum
Quantum yield, II photo-quantum efficiency of PS, Relative electron transport rate, Photochemical quenching coefficient and non-Photochemical quenching coefficient；With
Photosynthetic apparatus measures Net Photosynthetic Rate, transpiration rate, intercellular CO2Concentration, stomatal conductance；Choose the lower 3 plants of childrens of every group of light processing
The current year leave piece of seedling, is taken back in laboratory with ice chest, and every plant of blade is shredded mixing, green with ultraviolet specrophotometer measurement leaf
Plain a, b and carotenoid content.
8. a kind of method of measurement function group seedling Niche Differentiation on light resource axis according to claim 1, special
Sign is that the measuring method of level of resources utilization index includes: that the seedling leaves sample that will be fetched crushes after 70-90 DEG C of drying
Sieving, using the full nitrogen of Kjeldahl nitrogen determination；According to net primary production and the photosynthetically active radiation amount ratio absorbed
Calculate light utilization ratio, net photosynthetic rate and Dark respiration rate ratio calculate carbon utilization efficiency, Nitrogen productivity with
The product of nitrogen mean residence time calculates nitrogen use efficiency.
9. a kind of method of measurement function group seedling Niche Differentiation on light resource axis according to claim 1, special
Sign is, the measuring method of photosensitizing substance conversion ratio index includes: to react photosynthetic with Bomass allocation and blade assimilatory efficiency
Substance conversion ratio randomly selects several plants of seedling in every group of light processing, claims each organ to fresh weight respectively, and sample and take back experiment
Interior, 70-90 DEG C drying 36-60 hours, claim root, stem, leaf each section organ dry weight, convert amount of dry matter, and calculates following
Index: root again ratio, stem compare again, leaf mass ratio, ground/Underground biomass, Gao Jingchong when relative growth rate；In order to inquire into
The assimilatory efficiency difference of each level blade of tree crown, respectively weighs to each sequence side shoot, and current-year branch is individually taken
Sample weighing, using each organic unit's time biomass total amount in non-assimilation part, current year Leaf biomass and the total leaf area of individual as base
Plinth calculates the gross assimilation efficiency of blade, and the blade assimilatory efficiency of each organ is then calculated according to Bomass allocation ratio.
10. according to claim 1 to the side of any measurement function group seedling Niche Differentiation on light resource axis in 9
Method, which is characterized in that be the sample north subtropical Da Bie Mountain area Mixed Evergreen-deciduous Broad-leaved Forests.
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