CN108037240A - The plant division of one kind analysis clone plant, rhizome or stolon maximum keep alive aging method - Google Patents
The plant division of one kind analysis clone plant, rhizome or stolon maximum keep alive aging method Download PDFInfo
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- CN108037240A CN108037240A CN201711157154.1A CN201711157154A CN108037240A CN 108037240 A CN108037240 A CN 108037240A CN 201711157154 A CN201711157154 A CN 201711157154A CN 108037240 A CN108037240 A CN 108037240A
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- G01N—INVESTIGATING OR ANALYSING MATERIALS BY DETERMINING THEIR CHEMICAL OR PHYSICAL PROPERTIES
- G01N33/00—Investigating or analysing materials by specific methods not covered by groups G01N1/00 - G01N31/00
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Abstract
The present invention proposes a kind of plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method, and this method has key meaning for research rhizome type or stolon type clone plant species characteristic.
Description
Technical field
The present invention relates to phytoecology field, more particularly to a kind of plant division of analysis clone plant, rhizome or stolon are most
Big living age method.
Background technology
Analysis forms the individual age of plant population for understanding and predicting that population structure and dynamic are of great significance.Very
The age of more plants develops (including breeding, solid) character and/or pest and disease damage generation etc. with its growth substantial connection.For
For many economic plants, its age is had any actual knowledge of, it helps formulate rational management tactics.
Many biologies can form obvious natural trace in its growth growth course, therefore, analyze its age not
It is difficult.But the living age for analyzing rhizome type or stolon type clone plant is always in phytoecology field for a long time
Thorny problem, because general be difficult to discover whether they form other the similar lifes that can be used to the identification age in growth course
The natural trace of thing.The maximum keep alive age of rhizome type or stolon type clone plant is analyzed, for this kind of plant population,
With critical meaning.
The content of the invention
Studied by long-term, substantial amounts of field observation, sampling analysis, and greenhouse experiments, find many rhizome types or crawl
The age of crawl stem type clone plant can be identified by a kind of approach, and have feasibility.Based on this, the present invention proposes
A kind of plant division of analysis clone plant, rhizome or the method at stolon maximum keep alive age, comprise the following steps：
Step A：In the typical or representational habitat that specific rhizome type or stolon type clone plant are distributed, to it
Growth characteristics are investigated, and determine the Variation Features of its rhizome or each intercalary growth amount of stolon, plant division growth characteristic, plant division
Mean survival time (service life) and the variation characteristic of plant division form (size, the number of blade, biomass, leaf scar, branch trace, bud scale scar
Deng), particularly rhizome or stolon and plant division be in the unique adaptability spy shown during cyclically-varying in water, thermal resource
Sign, and rhizome or stolon and plant division be per annual total increment, as analysis and judges clone plant plant division, rhizome or crawls
The foundation of crawl stem living age；When rhizome or each intercalary growth amount of stolon are when water, thermal resource are in cyclically-varying, correspondingly
When showing cyclically-varying and/or other unique compliance characteristics, step B is gone successively to, otherwise terminates (this method discomfort
With other methods need to be used instead).
Step B：For specific rhizome type or stolon type clone plant, choose some long rhizomes or stolon is taken out
Inspection carries out continuing tracking (only focusing on the part of survival, exclude withered part)；
Step C：The first kind section and/or the second class section of each sampling observation rhizome of observation or stolon；The first kind
Section undergoes obvious periodicity rhythm and pace of moving things change at least once for the panel length and/or internode biomass of rhizome or stolon
Section；The second class section is the section with the imperfect rhythm and pace of moving things for cutting off rhizome or stolon when sampling and being formed, and/
Or growth cycle, one section of rhizome or stolon with apical growing point are not completed in sampling fashion；
According to the suitably Growing season of the plant or water, thermal resource seasonal variety number of cycles in 1 year, and rhizome or
The length of first section and the number of the periodicity rhythm and pace of moving things is included on stolon, calculate the curtate age of rhizome or stolon；
Each second section according to containing on rhizome or stolon accounts for the ratio of a complete rhythm and pace of moving things, estimates its growth institute's used time
Between, it is denoted as decimal age of rhizome or stolon；The cumulative decimal age is simultaneously added with curtate age, up to rhizome or stolon
Existent age；
According to observed in step A in each Growing season or when water, thermal resource seasonal variety the growth of plant division and
Morphological change characteristics and/or other unique compliance characteristics, with reference to the actual state of plant division, judge its growth to actual state
The required time, and the age is identified with this.
Step D：Collect, whole the sampling observation rhizomes or stolon and the age data of plant division that collation pass step C is obtained, push away
It is disconnected to obtain the average maximum keep alive age of specific clone plant plant division, rhizome or stolon.
It is proposed by the present invention to analyze in clone plant plant division, rhizome or stolon maximum keep alive aging method, the step
B, the clone plant directly observed for plant division, rhizome or stolon growing state inconvenience, is analyzed using quadrat method, it is wrapped
Include following steps：
Step B1：Determine to analyze the basic sample area and shape needed for its plant division, rhizome or stolon maximum keep alive age
Shape；
Step B2：Sample prescription is laid, along at least one side of the sample prescription, is sequentially arranged adjacent sample prescription；To across sample prescription
Specific the clone plant rhizome or stolon on border are implemented to cut off one by one；After cut-out, the script category to being respectively at notch both ends
In the segment of same rhizome or stolon, it is marked using identical label or numbering；
Step B3：To the specific clone plant in each sample prescription, if being not easy to directly observe, dig out and remove mud
Soil, according to above-mentioned steps A, B, C, D, observation is located at rhizome or stolon and the living age of plant division in sample prescription；If it is easy to
Directly observe, then do not dig out, now removing the soil for hindering observation, observation rhizome or stolon and the living age of plant division；
Step B4：In all adjacent serial sample prescriptions, according to label or the homogeney of numbering, select a plurality of across to the greatest extent may be used
Can how adjacent sample prescription rhizome or stolon sampling observation carry out continuing tracking, and collect the survival of each rhizome or stolon and plant division
Age.
It is proposed by the present invention to analyze in clone plant plant division, rhizome or stolon maximum keep alive aging method, the step
B, the clone plant for being easy to directly observe for plant division, rhizome or stolon growing state, using the method directly observed.
It is proposed by the present invention to analyze in clone plant plant division, rhizome or stolon maximum keep alive aging method, the root of sampling observation
Stem or stolon quantity are at least above 3, and the plant division quantity of sampling observation is at least above 3.
It is proposed by the present invention to analyze in clone plant plant division, rhizome or stolon maximum keep alive aging method, the root of sampling observation
The suitable number of stem or stolon is 30-50 bars (more persons are unlimited, the more better)；The suitable number for inspecting plant division by random samples is 30-50
(more persons are unlimited, the more better).
It is proposed by the present invention to analyze in clone plant plant division, rhizome or the method at stolon maximum keep alive age, sampling observation
Rhizome or stolon have following feature：The rhizome or stolon of same rank living, and originated from a ripe plant division living,
Extend in one direction growth, the panel length and/or internode biomass of rhizome or stolon with a Growing season periodicity
Change or water, the cyclically-varying experience obvious periodicity rhythm and pace of moving things change, and terminating at new life at least once of thermal resource
The apical growing point of internode and newborn plant division bud.
The plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method proposed by the present invention, for studying root
Stem type or stolon type clone plant species characteristic have key meaning.
Brief description of the drawings
Fig. 1 is to be used to analyze the two neighboring of clone plant plant division, rhizome or stolon maximum keep alive age in the present invention
The schematic diagram of subquadrat.
Embodiment
With reference to specific examples below and attached drawing, the invention will be described in further detail.Implement process, the bar of the present invention
Part, experimental method etc., in addition to the following content specially referred to, are among the general principles and common general knowledge in the art, the present invention
Content is not particularly limited.
In Fig. 1, crisscross line represents rhizome or stolon (thereon strain estranged), wherein a thick line is as generation
Table, represents that each section is shown in different Growing seasons or water, the growth in the thermal resource seasonal variety cycle with the segmentation of the different depths
Amount.These rhizomes or stolon are in originally attached state, by adjacent sample prescription, cut-out originally connected rhizome or stolon, it
Afterwards, it is marked with what the fragment of identical label or the numbering same rhizome or stolon connected to script was connected originally.It is small
Sample prescription can respectively be excavated, harvested, but the connection relation of rhizome or stolon can be followed the trail of continuously.
The present invention proposes a kind of plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method, including with
Lower step：
Step A：Select in the rhizome type of water, thermal resource in the area growth of obvious seasonal variety or stolon type clone
Plant is object.In the typical or representational habitat that specific rhizome type or stolon type clone plant are distributed, it is given birth to
Long characteristic is investigated, understand the Variation Features of its rhizome or each intercalary growth amount of stolon, plant division growth characteristic, plant division it is flat
Equal time-to-live (service life) and the variation characteristic (size, the number of blade, biomass, leaf scar, branch trace, bud scale scar etc.) of plant division form,
Or its other unique compliance characteristics for showing in water, thermal resource change, and rhizome or stolon and plant division it is annual
Average total increment a etc., as the foundation for analyzing and judging clone plant plant division, rhizome or stolon living age.
Step B：According to the Variation Features and development of the rhizome of specific clone plant or each intercalary growth amount of stolon
The comfort level of actual observation, determines to analyze its plant division, rhizome or the method at stolon maximum keep alive age：Method of direct observation and
Quadrat method.
Specifically, method of direct observation：Being easy to directly observation for plant division, rhizome or stolon growing state (need not break
Rhizome or stolon are cut off or excavated to bad property) clone plant, method of direct observation can be used：Choose some long rhizomes or
Stolon CI(CIRepresenting selected specific rhizome or stolon) (at least above 3,30-50 bars are advisable) sampling observation held
Continuous tracking (only focusing on the part of survival, exclude withered part).Inspect C by random samplesI(CIImplication is same as above) there need to be following essential characteristic：
The rhizome or stolon of same rank living, and from a ripe plant division starting living, extends in one direction growth, rhizome or
The panel length of stolon and/or internode biomass become with the cyclically-varying or water, the periodicity of thermal resource of Growing season
Change experience obvious periodicity rhythm and pace of moving things change at least once, and terminate at the apical growth with newborn internode and newborn plant division bud
Point.
Specifically, quadrat method：(destructiveness is needed to cut for plant division, rhizome or stolon growing state inconvenience directly observation
Disconnected or excavate rhizome or stolon, could analyze) clone plant, quadrat method can be used：Determine analyze its plant division, rhizome or
Basic sample area b needed for the stolon maximum keep alive ageiWith shape (biRepresent the growth characteristics according to specific clone plant
The Sampling Area determined with actual habitat i).When sequentially continuing sampling, using a line as boundary between adjacent sample prescription, fit closely.
In conditions permit, biIt is preferably as larger as possible, in order to being tracked observation in continuously distributed big clone plant.
In the typical or representational habitat that specific rhizome type or stolon type clone plant are distributed, sample prescription is laid,
To across sample prescription bi(biImplication is same as above) specific the clone plant rhizome or stolon c on borderi(ciRepresent specific certain rhizome or
Stolon) implement cut-out one by one.Cutting off every ci(ciImplication is same as above) after, immediately to notch both ends originally belong to it is same
The fragment of rhizome or stolon, is marked using identical label or numbering, for same in adjacent sample prescription to being distributed in
Rhizome or stolon carry out continuing tracking.According to the comfort level of field observation, select to sample prescription bi(biImplication is same as above) in tool
Body clone plant, if being not easy to directly observe, digs out and removes soil, and according to above-mentioned steps A, B, C, D, observation is located at sample
The living age of rhizome or stolon and plant division in side；If easy to directly observe, do not dig out, now removing obstruction sight
The soil of survey, observe rhizome or stolon and the living age of plant division.
In order to being tracked observation in continuously distributed big clone plant, along sample prescription bi(biImplication is same as above) at least
One side, is sequentially arranged adjacent sample prescription bi+1(biImplication is same as above), by the method for step 3, to sample prescription bi+1(biImplication is same as above) into
Row processing；And so on, to sample prescription bi+j(biImplication is same as above, and j represents sample prescription bi(biImplication is same as above) after adjacent jth successively
A sample prescription) handled.
In all adjacent serial sample prescriptions, according to label or the homogeney of numbering, select a plurality of across multiphase as far as possible
The rhizome or stolon C of adjacent sample prescriptionI(CIRepresent selected specific rhizome or stolon) (at least above 3,30-50 bars are
Preferably, more persons are unlimited, the more better) sampling observation carry out continue tracking.Inspect C by random samplesI(CIImplication is same as above) there need to be following essential characteristic：It is living
Same rank rhizome or stolon, and from a ripe plant division starting living, extend in one direction growth, rhizome or crawl
The panel length of crawl stem and/or internode biomass with a Growing season cyclically-varying or water, the cyclically-varying of thermal resource
Experience obvious periodicity rhythm and pace of moving things change at least once, and terminate at the apical growth with newborn internode and newborn plant division bud
Point.
Step C：Rhizome or stolon C for sampling observationI(CIImplication is same as above), observe each CI(CIImplication is same as above) it is every
A panel length (preferably while surveying biomass).The rhizome or stolon C of sampling observationI(CIImplication is same as above) two categories below may be included
Section (or wherein first kind section)：First kind section R, panel length and/or internode biomass with a Growing season cycle
Property change or water, thermal resource multiple (at least once) the obvious periodicity rhythm and pace of moving things r changes of cyclically-varying experience section；The
Class two zone section Δ R, the section with imperfect rhythm and pace of moving things r for cutting off rhizome or stolon when sampling and being formed, and/or sampling
Fashion does not complete growth cycle and r's and terminates at one section of rhizome or stolon with apical growing point.
Rhizome or stolon CI(CIImplication is same as above) on, it is a life the time required to each complete periodicity rhythm and pace of moving things r growths
Long season or water, the cycle of thermal resource seasonal variety.According to the suitably Growing season of the clone plant or water, thermal resource season in 1 year
Section property period of change number, and rhizome or stolon CI(CIImplication is same as above) length of upper curtate R and the number comprising r, can be with
Calculate rhizome or stolon CI(CIImplication is same as above) curtate age Wi(i represents specific certain rhizome or stolon).And for root
Stem or stolon CI(CIImplication is same as above) on Δ Zone R section, the ratio of a complete rhythm and pace of moving things r is accounted for according to every section of Δ R, can be estimated
Time used in its growth, and it is denoted as decimal age wi(wiRepresent the time used in specific certain rhizome or the growth of stolon fragment).It is tired
Add wiAnd with curtate age Wi(i represents specific certain rhizome or stolon) is added, up to rhizome or stolon CI(CIImplication is same
On) living age.
The each section of observation is gone up raw plant division quantity (preferably while measure biomass), is observed according in step A every
The growth of plant division and morphological change characteristics in a Growing season or when water, thermal resource seasonal variety (size, the number of blade, biomass,
Leaf scar, branch trace, bud scale scar when) and/or its in water, thermal resource change when other unique compliance characteristics for showing, with reference to
The actual state of plant division, judges the time needed for its growth to actual state, and identifies the age with this.
Step D：Collect, whole the sampling observation rhizomes or stolon C that collation pass step C is obtainedI(CIImplication is same as above) and plant division
Age data, deduction obtain the average maximum keep alive age of specific clone plant plant division, rhizome or stolon.
In the present invention, when in the serial sample prescription, the rhizome or stolon C of sampling observationI(CIImplication is same as above) quantity is not up to pre-
During fixed number amount (at least above 3,30-50 bars are advisable, and more persons are unlimited, the more better), then it is distributed in specific clone plant
Typical or representational habitat continue to set, select and observe, until meet predetermined quantity (at least above 3,
30-50 bars are advisable, and more persons are unlimited, the more better).
The protection content of the present invention is not limited to above example.Without departing from the spirit and scope of the invention, originally
Field technology personnel it is conceivable that change and advantage be all included in the present invention, and using appended claims as protect
Protect scope.
Claims (7)
1. the plant division of one kind analysis clone plant, rhizome or stolon maximum keep alive aging method, it is characterised in that including following step
Suddenly：
Step A：In the typical or representational habitat that specific rhizome type or stolon type clone plant are distributed, to its growth
Characteristic is investigated, and determines that the Variation Features of its rhizome or each intercalary growth amount of stolon, plant division growth characteristic, plant division are averaged
Time-to-live and the variation characteristic of plant division form, particularly rhizome or stolon and plant division are in cyclically-varying in water, thermal resource
When unique compliance characteristics for showing, and rhizome or stolon and plant division are per annual total increment；When rhizome or crawl
Each intercalary growth amount of crawl stem correspondingly shows the suitable of cyclically-varying and/or uniqueness when water, thermal resource are in cyclically-varying
During answering property feature, step B is gone successively to, is otherwise terminated；
Step B：For specific rhizome type or stolon type clone plant, choose some long rhizomes or stolon inspect by random samples into
The lasting tracking of row, wherein, the part of survival is only focused on, excludes withered part；
Step C：The first kind section and/or the second class section of each sampling observation rhizome of observation or stolon；The first kind section
For the panel length and/or internode biomass of rhizome or stolon the experience area that the obvious periodicity rhythm and pace of moving things changes at least once
Section；The second class section is the section with the imperfect rhythm and pace of moving things for cutting off rhizome or stolon when sampling and being formed, and/or
Growth cycle, one section of rhizome or stolon with apical growing point are not completed in sampling fashion；
According to the suitably Growing season of the plant or water, thermal resource seasonal variety number of cycles in 1 year, and rhizome or crawl
The length of first section and the number of the periodicity rhythm and pace of moving things is included on stem, calculate the curtate age of rhizome or stolon；According to
Each second section contained on rhizome or stolon accounts for the ratio of a complete rhythm and pace of moving things, estimates the time used in its growth,
It is denoted as the decimal age of rhizome or stolon；The cumulative decimal age is simultaneously added with curtate age, up to the life of rhizome or stolon
Deposit the age；
According to being observed in step A in each Growing season or the growth of plant division and form when water, thermal resource seasonal variety
Variation characteristic and/or other unique compliance characteristics, with reference to the actual state of plant division, judge needed for its growth to actual state
Time, and the age is identified with this；
Step D：Collect, whole the sampling observation rhizomes or stolon and the age data of plant division that collation pass step C is obtained, infer
To the average maximum keep alive age of specific clone plant plant division, rhizome or stolon.
2. the plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method, its feature exist as claimed in claim 1
In in the step A, the variation characteristic of the plant division form includes size, the number of blade, biomass, leaf scar, branch trace, bud scale scar.
3. the plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method, its feature exist as claimed in claim 1
In, in the step B, the clone plant directly observed for plant division, rhizome or stolon growing state inconvenience, using quadrat method
Analyzed, it comprises the following steps：
Step B1：Determine to analyze the basic sample area and shape needed for its plant division, rhizome or stolon maximum keep alive age；
Step B2：Sample prescription is laid, along at least one side of the sample prescription, is sequentially arranged adjacent sample prescription；To across sample prescription border
Specific clone plant rhizome or stolon implement to cut off one by one；After cut-out, to being respectively at originally belonging to together for notch both ends
The segment of one rhizome or stolon, is marked using identical label or numbering；
Step B3：To the specific clone plant in each sample prescription, if being not easy to directly observe, dig out and remove soil, press
According to described step A, B, C, D, observation is located at rhizome or stolon and the living age of plant division in sample prescription；If easy to directly see
Survey, then do not dig out, now removing the soil for hindering observation, observation rhizome or stolon and the living age of plant division；
Step B4：In all adjacent serial sample prescriptions, according to label or the homogeney of numbering, select a plurality of across as more as possible
Rhizome or the stolon sampling observation of adjacent sample prescription carry out continuing tracking, and collect the survival year of each rhizome or stolon and plant division
Age.
4. the plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method, its feature exist as claimed in claim 1
In, in the step B, the clone plant that is easy to directly observe for plant division, rhizome or stolon growing state, using directly seeing
The method of survey.
5. the plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method, its feature exist as claimed in claim 1
In the rhizome or stolon quantity of sampling observation are at least above 3, and the plant division quantity of sampling observation is at least above 3.
6. the plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method, its feature exist as claimed in claim 1
In the rhizome of sampling observation or the suitable number of stolon are at least 30；The suitable number for inspecting plant division by random samples is at least 30.
7. the plant division of analysis clone plant, rhizome or stolon maximum keep alive aging method, its feature exist as claimed in claim 1
In the rhizome or stolon of sampling observation have following feature：The rhizome or stolon of same rank living, and from a maturation living
Plant division originates, and extends in one direction growth, the panel length and/or internode biomass of rhizome or stolon are with a Growing season
Cyclically-varying or water, the cyclically-varying experience obvious periodicity rhythm and pace of moving things change, and terminating at least once of thermal resource
Apical growing point with newborn internode and newborn plant division bud.
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| CN113960245A (en) * | 2021-09-02 | 2022-01-21 | 华东师范大学 | Approximate determination method for contribution of clone integration to internode formation of rhizome type or stolon type clone plant |
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| CN113780623A (en) * | 2021-08-09 | 2021-12-10 | 华东师范大学 | Method for differentially analyzing age structure of herbaceous clone plant division population and age structure of rootstock or stolon supported by herbaceous clone plant division population |
| CN113960245A (en) * | 2021-09-02 | 2022-01-21 | 华东师范大学 | Approximate determination method for contribution of clone integration to internode formation of rhizome type or stolon type clone plant |
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