Latin name of the genus and species of the plant claimed: Rubus idaeus.
Variety denomination: ‘ENDURANCE’.
BACKGROUND
The present invention relates to a new and distinct everbearing raspberry variety designated ‘ENDURANCE’. This new variety is a result of a controlled cross made in 2008 in Watsonville, Calif. between raspberry variety ‘GRANDEUR’ (patented, U.S. Plant Pat. No. 20,459) as the female parent and raspberry variety ‘04.3891’ (unpatented) as the male parent in an ongoing breeding program. The variety is botanically known as Rubus idaeus and was tested as ‘PS-08.056-18’ and ‘PS-5618’.
The seedling resulting from the aforementioned cross was asexually propagated by dormant canes in Santa Cruz County, Calif. and was subsequently selected by the inventor from a controlled breeding plot in Watsonville, Calif. in 2010. After its selection, the new variety was further asexually propagated by dormant canes, roots and non-dormant root shoot cuttings in Santa Cruz County, Calif., San Joaquin County, Calif. and Siskiyou County, Calif. The new variety was then extensively tested over the next several years in fruiting fields in Santa Cruz County, Calif. and has performed successfully in grower fields in USDA zones 9-10. This propagation has demonstrated that the combination of traits disclosed herein as characterizing the new variety are fixed and remain true to type through successive generations of asexual reproduction.
SUMMARY
‘ENDURANCE’ is primarily adapted to the climate and growing conditions of the central coast of California. This region provides the necessary year-round temperatures required for it to produce and maintain a strong vigorous plant and to remain in fruit production from July through December on primocanes and in the ensuing year from May through July on the floricanes. The following traits have been repeatedly observed and are determined to be unique characteristics of ‘ENDURANCE’, which in combination distinguish this raspberry plant as a new and distinct variety:
-
- 1. Dark purple spines
- 2. Late fruit yield
- 3. Medium red fruit color
- 4. Absent to very weak primocane glaucosity
- 5. Fair flavor
The raspberry variety that is believed to be most closely related to the new raspberry variety ‘ENDURANCE’ is the raspberry variety ‘GRANDEUR’ (patented, U.S. Plant Pat. No. 20,459). In comparison to the similar raspberry variety ‘GRANDEUR’, ‘ENDURANCE’ differs by the following combination of characteristics described in Table 1:
COMPARISON BETWEEN ENDURANCE AND GRANDEUR
| TABLE 1 |
| |
| |
|
’GRANDEUR’ |
| Characteristic |
‘ENDURANCE’ |
(US PP20,459) |
| |
| Primocane waxy |
Absent to very |
Absent to very |
| coat (glaucosity) |
weak |
weak |
| Predominate number |
Equal 3-5 |
Always 3 |
| of leaflets |
|
|
| Leaf shape |
Flat to slightly |
Flat to slightly |
| (cross section) |
convex |
convex |
| Rugosity |
Weak |
Strong |
| Color mature fruit |
RHS 43A |
RHS 42A |
| |
Medium red |
Medium red |
| Thorn Tip Coloration |
N186B |
180C |
| |
Greyed-purple |
Greyed-red |
| Relative number of |
26.9 |
16.2 |
| pedicel thorns |
Many |
Medium |
| |
‘ENDURANCE’ differs from its sibling raspberry variety ‘MAJESTIC’ (patented, U.S. Plant Pat. No. 31,716) by the following combination of characteristics described in Table 2:
COMPARISON BETWEEN ENDURANCE AND MAJESTIC
| TABLE 2 |
| |
| |
|
‘MAJESTIC’ |
| |
|
(U.S. Plant Pat. |
| Characteristic |
‘ENDURANCE’ |
No. 31,716) |
| |
| Time of fruiting (primocane) |
Late |
Medium |
| Terminal leaf shape |
Lobed |
Cordate |
| Leaf shape (cross section) |
Flat to slightly convex |
Slightly convex |
| Rugosity |
Weak |
Medium |
| Relative size of drupes |
Large |
Medium |
| Glossiness (fruit) |
Medium |
High |
| Adherence of receptacle |
Very weak |
Weak |
| Flavor |
Fair |
Very good |
| |
‘ENDURANCE’ differs from its parents, ‘GRANDEUR’ and ‘04.3891’, by the following combination of characteristics described in Table 3:
COMPARISON BETWEEN ENDURANCE, GRANDEUR AND 04.3891
| TABLE 3 |
| |
| |
|
‘GRANDEUR’ |
|
| |
|
(U.S. Plant Pat. |
|
| Characteristic |
‘ENDURANCE’ |
No. 20,459) |
‘04.3891’ |
| |
| Productivity |
High |
High |
Medium |
| Glossiness (fruit) |
Medium |
Medium |
High |
| Firmness (fruit) |
Medium to high |
Very high |
Medium to low |
| Adherence of |
Very weak |
Very weak |
Medium |
| receptacle |
|
|
|
| Primocane time |
Late |
Medium |
Medium |
| of fruiting |
|
|
|
| Fruit size |
Medium to large |
Medium |
Large |
| Flavor |
Fair |
Good |
Good |
| |
For identification, a series of molecular markers have been determined for this new variety.
BRIEF DESCRIPTION OF THE DRAWINGS
The accompanying color photographs illustrate the overall appearance of typical specimens of the new raspberry variety ‘ENDURANCE’ at various stages of development as true as reasonably possible with color reproductions of this type. Color in the photographs may differ slightly from the color value cited in the detailed botanical description which accurately describes the color of ‘ENDURANCE’. The depicted plant and plant parts of the new raspberry variety ‘ENDURANCE’ were taken in Watsonville, Calif. and are approximately 2 to 16 months old:
FIG. 1 shows typical leaf shape, foliate and rugosity characteristics of ‘ENDURANCE’ taken in the month of September 2020;
FIG. 2 shows bud, flower and fruit characteristics of ‘ENDURANCE’ taken in the month of September 2020;
FIG. 3 shows typical gloss, coloration, and drupelet formation of primocane fruit of ‘ENDURANCE’ taken in the month of September 2020;
FIG. 4 shows typical primocane and spine coloration of ‘ENDURANCE’ taken in the month of September 2020;
FIG. 5 shows typical dormant bud characteristics of ‘ENDURANCE’ taken in the month of January 2019.
DETAILED BOTANICAL DESCRIPTION
‘ENDURANCE’ has not been observed under all possible environmental conditions. The characteristics of the new variety may vary in detail, depending upon variations in environmental factors, including weather (temperature, humidity and light intensity), day length, soil type, and location.
The aforementioned photographs, together with the following description of the new raspberry variety ‘ENDURANCE’, unless otherwise noted, are based upon observations taken during the 2016-2017 growing season in Watsonville, Calif. Primocane measurements and ratings were taken from plants of ‘ENDURANCE’ dug from a nursery located in Siskiyou County, Calif. during the middle of October 2015 and planted approximately 3 to 4 weeks later in Watsonville, Calif. The approximate age of the observed primocane plants is 7 to 8 months. Floricane measurements and ratings were taken from the same planting of ‘ENDURANCE’ at an approximate age of 16 to 18 months. Yield observations and fruit quality characteristics are averaged from 3 years of data collected from the 2015 through 2018 production seasons. Flower measurements and characteristics are from secondary flowers unless otherwise noted. Fruit characteristics and measurements are from secondary fruit unless otherwise noted. Foliage characteristics and measurements are from 3-foliate foliage unless otherwise noted.
Color terminology where noted follows The R.H.S. Colour Chart Fifth Edition, Royal Horticultural Society, London, United Kingdom (2007).
The following tables 3-7 describe fruit, plant, foliage, flower and pest/disease characteristics of the new raspberry ‘ENDURANCE’ in comparison to the similar raspberry varieties ‘GRANDEUR’ (patented, U.S. Plant Pat. No. 20,459).
COMPARISON BETWEEN ENDURANCE AND GRANDEUR
| TABLE 3 |
| |
| FRUIT CHARACTERISTICS |
| |
|
|
‘GRANDEUR’ |
| |
|
|
(U.S. Plant Pat. |
| |
Characteristic |
‘ENDURANCE’ |
No. 20,459) |
| |
| |
Color mature fruit |
RHS 42A |
RHS 42A |
| |
|
Medium red |
Medium red |
| |
Color achenes |
RHS 159A |
RHS 159A |
| |
|
Orange-white |
Orange-white |
| |
|
group |
group |
| |
Fruit length (mm) |
27.71 |
22.91 |
| |
Fruit width (mm) |
23.20 |
20.12 |
| |
Length/Width ratio |
1.19 |
1.14 |
| |
|
Longer than |
Longer than |
| |
|
broad |
broad |
| |
Seed weight (mg) |
1.76 |
1.75 |
| |
Drupelets per berry |
95 |
93 |
| |
Weight of single drupe |
0.071 |
0.043 |
| |
(g/drupe) |
|
|
| |
Drupe length (mm) |
7.45 |
5.25 |
| |
Drupe diameter (mm) |
4.53 |
3.86 |
| |
Relative size of drupes |
Large |
Medium |
| |
Fruit size |
Medium |
Medium |
| |
Fruit per node |
10 |
15 |
| |
Predominant shape |
Conical |
Conical |
| |
Receptacle length (mm) |
24.74 |
19.15 |
| |
Receptacle diameter (mm) |
12.02 |
10.32 |
| |
Receptacle color |
RHS 155B |
RHS 155B |
| |
|
White group |
White group |
| |
Evenness of color (fruit) |
Even |
Even |
| |
Glossiness |
Medium |
Medium |
| |
Adherence of receptacle |
Very weak |
Very weak |
| |
Firmness of flesh |
Firm |
Very firm |
| |
Firmness of skin |
Firm |
Very firm |
| |
Soluble Solids (% brix) |
9.8 |
10.5 |
| |
Flavor |
Fair |
Good |
| |
Shipping quality |
Good |
Good |
| |
Fruit storage life (avg) |
9 days |
12 days |
| |
Fruit market use |
Fresh |
Fresh |
| |
| TABLE 4 |
| |
| PLANT CHARACTERISTICS |
| |
|
‘GRANDEUR’ |
| |
|
(U.S. Plant Pat. |
| Characteristic |
‘ENDURANCE’ |
No. 20,459) |
| |
| General: |
|
|
| Habit |
Semi-erect |
Upright |
| Plant height (m) |
1.5 |
1.4 |
| Plant width (cm) |
38 |
45 |
| Time to initiate roots (days) |
12-14 |
12-14 |
| Canes per hill |
7.5 |
6.9 |
| Productivity |
High |
High |
| Self-fruitfulness |
Yes |
Yes |
| Type of bearing |
Everbearing |
Everbearing |
| Primocane: |
|
|
| Color (true) |
RHS 145A |
RHS 145B |
| |
Yellow-green |
Yellow-green |
| |
group |
group |
| Length (cm) |
153.5 |
143.6 |
| Basal diameter (mm) |
19.26 |
21.80 |
| Diameter central ⅓ (mm) |
13.76 |
12.55 |
| Lateral length at central ⅓ (cm) |
27.7 |
43.6 |
| No. fruiting laterals per cane |
11.8 |
17.4 |
| Total nodes per cane |
39 |
31 |
| Internode length at central ⅓ (mm) |
38.93 |
48.84 |
| Anthocyanin coloration |
RHS 59B |
RHS 59B |
| |
Red-purple |
Red-purple |
| |
group |
group |
| Anthocyanin intensity |
Medium |
Weak |
| Pubescence |
Absent |
Absent |
| Vegetative bud length (mm) |
8.70 |
11.01 |
| Vegetative bud diameter (mm) |
5.15 |
6.11 |
| Vegetative bud shape |
Widely ovate |
Widely ovate |
| Vegetative bud color |
RHS 187A |
RHS 187A |
| |
Greyed-red |
Greyed-red |
| |
group |
group |
| Strength of waxy coat (glaucosity) |
Absent to very |
Absent to very |
| |
weak |
weak |
| Time of cane emergence |
Medium to late |
Medium |
| Time of flowering |
Medium to late |
Medium |
| Time of fruiting |
Medium to late |
Medium |
| Length of fruiting season |
Long |
Long |
| % of total yield |
38% |
50% |
| Flowering period |
Late June to |
Late June to |
| |
Late November |
Late November |
| Harvest period |
Late July to |
Late July to |
| |
Late December |
Late December |
| Primocane fruit weight (g) |
4.7 |
3.8 |
| Primocane yield (g/plant) |
1,936 |
2,662 |
| Young Shoots: |
|
|
| Number (per meter) |
15-20 |
15-20 |
| |
Medium |
Medium |
| Anthocyanin presence |
Present |
Present |
| Anthocyanin coloration |
RHS 179A |
RHS 179A |
| |
Red-purple |
Greyed-red |
| |
group |
group |
| Anthocyanin intensity |
Weak |
Medium |
| Thorns: |
|
|
| Thorn presence |
Present |
Present |
| Thorn coloration (tip) |
RHS 187A |
RHS 180C |
| |
Greyed-purple |
Greyed-red |
| |
group |
group |
| Thorn coloration (base) |
RHS N186C |
RHS 145B |
| |
Greyed-purple |
Yellow-green |
| |
group |
group |
| Thorn length at central ⅓ (mm) |
1.39 |
0.76 |
| Thorn base at central ⅓ (mm) |
1.32 |
1.53 |
| Thorn density per cm at central ⅓ |
5.25 |
3.38 |
| |
Medium |
Medium |
| Thorn texture |
Rigid |
Rigid |
| Attitude of the tip |
Horizontal |
Horizontal |
| Floricane: |
|
|
| Color (true) |
RHS 165B |
RHS 164A |
| |
Greyed-orange |
Greyed-orange |
| |
group |
group |
| Length (cm) |
123.4 |
119.8 |
| Width at central ⅓ (mm) |
12.58 |
12.04 |
| Total nodes per cane |
22 |
18 |
| Internode length at central ⅓ (mm) |
44.55 |
49.13 |
| Fruiting lateral attitude |
Erect |
Erect |
| Time bud burst (vegetative) |
Medium |
Medium |
| Time of flowering |
Medium |
Medium |
| Time of fruiting |
Medium |
Medium |
| Length of fruiting season |
Medium to long |
Medium to long |
| % of total yield |
62% |
50% |
| Flowering period |
Late April to |
Late April to |
| |
Late June |
Late June |
| Harvest period |
Late May to |
Late May to |
| |
Late July |
Late July |
| Floricane fruit weight (g) |
4.6 |
3.7 |
| Floricane yield (g/plant) |
3,161 |
2,680 |
| |
| TABLE 5 |
| |
| FOLIAGE CHARACTERISTICS |
| |
|
‘GRANDEUR’ |
| |
|
(U.S. Plant Pat. |
| |
‘ENDURANCE’ |
No. 20,459) |
| Characteristic |
(3 Foliate) |
(3 Foliate) |
| |
| General: |
|
|
| Color of upper surface |
RHS N137A |
RHS N137A |
| |
Green group |
Green group |
| Color of lower surface |
RHS 190B |
RHS 190B |
| |
Greyed-green |
Greyed-green |
| |
group |
group |
| Venation |
Pinnate |
Pinnate |
| Shape in cross section |
Flat to slightly |
Flat to slightly |
| |
convex |
convex |
| Arrangement |
Compound |
Compound |
| Relief between veins |
Weak |
Strong |
| (rugosity) |
|
|
| Glossiness |
Weak |
Weak |
| Number of leaflets/leaf |
Equal 3-5 |
Always 3 |
| Total leaf length (cm) |
26.1 |
22.5 |
| Total leaf width (cm) |
24.2 |
23.4 |
| Terminal Leaflet: |
|
|
| Length (mm) |
165.1 |
127.5 |
| Width (mm) |
120.3 |
91.6 |
| Length/width ratio |
1.4 |
1.4 |
| |
Longer than |
Longer than |
| |
broad |
broad |
| Size |
Large |
Medium to large |
| Shape |
Lobed |
Cordate |
| Shape of base |
Cordate |
Cordate |
| Shape of tip |
Acuminate |
Acuminate |
| Margins |
Biserrate |
Biserrate |
| Lateral Leaflet: |
|
|
| Length (mm) |
128.3 |
105.5 |
| Width (mm) |
74.1 |
65.1 |
| Length/width ratio |
1.7 |
1.6 |
| |
Much longer |
Longer than broad |
| |
than broad |
|
| Rachis length (mm) |
41.3 |
36.2 |
| Rachis diameter (mm) |
2.41 |
1.92 |
| Rachis anthocyanin coloration |
RHS 184B |
RHS 184A |
| of upper surface |
|
|
| Rachis anthocyanin intensity |
Weak |
Medium |
| Orientation |
Opposite |
Opposite |
| Arrangement |
Compound |
Compound |
| Relative position of lateral |
Free |
Touching |
| leaflets |
|
|
| Shape |
Ovate |
Ovate |
| Shape of the base |
Rounded |
Oblique rounded |
| Shape of the tip |
Acuminate |
Acuminate |
| Margins |
Biserrate |
Biserrate |
| Petiole: |
|
|
| Length (mm) |
54.54 |
59.00 |
| Width (mm) |
3.50 |
3.99 |
| Thorn presence |
Yes |
Yes |
| Thorn orientation |
Erect |
Erect |
| Anthocyanin coloration |
RHS 180B |
RHS 184C |
| of upper surface |
Greyed-purple |
Greyed-purple |
| |
group |
group |
| Anthocyanin intensity |
Absent to very |
Weak |
| |
weak |
|
| of upper surface |
|
|
| Stipule length (mm) |
12.77 |
10.23 |
| Stipule orientation |
Erect |
Erect to horizontal |
| |
| TABLE 6 |
| |
| FLOWER CHARACTERISTICS |
| |
|
‘GRANDEUR’ |
| |
|
(U.S. Plant Pat. |
| Characteristic |
‘ENDURANCE’ |
No. 20,459) |
| |
| Petal color (upper surface) |
RHS 155C |
RHS 155C |
| |
White group |
White group |
| Petal color (lower surface) |
RHS 155C |
RHS 155C |
| |
White group |
White group |
| Flower diameter (mm) |
27.25 |
22.55 |
| Petal length (mm) |
8.25 |
6.42 |
| Petal width (mm) |
3.42 |
3.11 |
| Petal length/width ratio |
2.41 |
2.06 |
| |
Much longer |
Much longer |
| |
than broad |
than broad |
| Petal shape in cross section |
Flat |
Flat |
| Petal shape (overall) |
Ovate |
Ovate |
| Petal shape (apex) |
Rounded |
Rounded |
| Petal shape (base) |
Attenuate |
Attenuate |
| Petal margin |
Entire |
Entire |
| Petal texture |
Glabrous |
Glabrous |
| No. petals/flower |
5.2 |
5.2 |
| No. sepals/flower |
5.2 |
5.2 |
| Sepal length (mm) |
10.80 |
11.20 |
| Sepal width at base (mm) |
4.41 |
6.49 |
| Sepal shape |
Deltate |
Widely deltate |
| Sepal shape (apex) |
Acuminate |
Acuminate |
| Sepal coloration (upper surface) |
RHS 144A |
RHS 144B |
| Sepal coloration (lower surface) |
RHS 144B |
RHS 144C |
| Sepal margin |
Entire |
Entire |
| Sepal texture |
Slightly |
Slightly |
| |
pubescent |
pubescent |
| No. stamen/flower |
116 |
105 |
| Filament length (mm) |
5.19 |
4.45 |
| Filament color |
RHS 155C |
RHS 155C |
| |
White group |
White group |
| Pollen quantity |
Medium |
Medium |
| Pollen color |
RHS 196D |
RHS 196D |
| Anther length (mm) |
1.16 |
0.79 |
| Anther diameter (mm) |
0.53 |
0.49 |
| Anther coloration |
RHS 155C |
RHS 155C |
| (pre-dehiscence) |
|
|
| Anther coloration |
RHS 165B |
RHS 165B |
| (post-dehiscence) |
|
|
| Stigma shape |
Lobed |
Lobed |
| Stigma length (mm) |
0.12 |
0.10 |
| Stigma diameter (mm) |
0.42 |
0.40 |
| Stigma coloration |
RHS 145D |
RHS 145D |
| Style length (mm) |
3.08 |
2.03 |
| Style diameter (mm) |
0.40 |
0.39 |
| Style coloration |
RHS 145D |
RHS 145D |
| Ovule length (mm) |
1.56 |
1.51 |
| Ovule diameter (mm) |
1.47 |
1.40 |
| Ovule coloration |
RHS 145B |
RHS 145B |
| Pedicle length (mm) |
21.55 |
23.86 |
| Pedicle diameter (mm) |
1.51 |
1.47 |
| Pedicle anthocyanin coloration |
RHS 181C |
RHS 178A |
| Pedicle anthocyanin intensity |
Absent to very |
Medium |
| |
weak |
|
| Relative number of pedicel |
26.9 |
16.2 |
| thorns |
Many |
Medium |
| Peduncle anthocyanin presence |
Present |
Present |
| Peduncle anthocyanin coloration |
RHS 184C |
RHS 184A |
| |
Greyed-purple |
Greyed-purple |
| |
group |
group |
| Peduncle anthocyanin intensity |
Weak |
Medium |
| |
| TABLE 7 |
| |
| PEST AND DISEASE REACTIONS |
| |
|
‘GRANDEUR’ |
| |
|
(U.S. Plant Pat. |
| Characteristic |
‘ENDURANCE’ |
No. 20,459) |
| |
| Spotted wing drosophila |
Susceptible |
Susceptible |
| (Drosophila suzukii) |
|
|
| Two spotted spider mite |
Susceptible |
Susceptible |
| (Tetranychus urticae) |
|
|
| Grey fruit mold |
Susceptible |
Susceptible |
| (Botrytis cinerea) |
|
|
| Powdery mildew |
Moderately |
Moderately |
| (Podosphaera aphanis var. aphanis) |
resistant |
susceptible |
| Yellow rust |
Moderately |
Moderately |
| (Phragmidium rubi-idaei) |
resistant |
susceptible |
| |